|Scientific Name:||Ursus arctos|
|Species Authority:||Linnaeus, 1758|
|Red List Category & Criteria:||Least Concern ver 3.1|
|Assessor(s):||McLellan, B.N., Proctor, M.F., Huber, D. & Michel, S.|
|Contributor(s):||Ambarli, H., Ararat, K., Aryal, A., Badamjav, L., Batmunkh, M., Can, O., Davletbakov, A., Esipov, A., Galbreath, G., Ghaemi, R., Gong, J., Gutleb, B., Han, S., Harris, R., Kaczensky, P., Kubanichbek, J., Lortkipanidze, B., Lukarevskiy, V., Mano, T., Moheb, Z., Nawaz, M.A., Oromov, B., Paczkowski, J., Puchkovskiy, S., Reynolds, H., Saidov, A., Sathyakumar, S., Sato, Y., Seryodkin, I., Tserenbataa, T., Tsuruga, H., Vaisfeld, M. & Xu, A.|
The range of the Brown Bear has historically declined in North America, Europe, and Asia, and the species has been extirpated in North Africa. However, it remains widespread across three continents, and is still one of the world’s most widely distributed terrestrial mammals. Globally the population remains large, and is not significantly declining and may be increasing in some areas (Swenson et al. 1998, Schwartz et al. 2006, Mace et al. 2012, Kaczensky et al. 2013, Chapron et al. 2014). There are many small, isolated populations that are in jeopardy of extirpation, but others, under more protection and management, are expanding.
|Previously published Red List assessments:|
The Brown Bear is the most widely distributed ursid. It once ranged across a large portion of western North America, including northern Mexico. There is also some evidence of sporadic occurrence (after retreat of the Wisconsin ice sheet) within the eastern half of the continent (Guilday 1968). Populations on the Ungava Peninsula may have extended to the Atlantic Ocean (Loring and Spiess 2007). They also ranged throughout Europe, Asia, the Middle East, and across North Africa. A history of prolonged overexploitation in Europe stretching back centuries resulted in the elimination of brown bears from many countries. The date of their extirpation from North Africa is uncertain, but they may have existed as late as the 1500s in the Sinai of Egypt (Manlius 1998) and mid-1800s in Algeria and Morocco (Hamdine et al. 1998). During the 20th century, Brown Bears (called Grizzly Bears in interior North America) were intentionally extirpated in Mexico (≈1960), a large portion of southwestern U.S. (Brown 1985, Mattson and Merrill 2002), and most of the Canadian prairies. Bears have been rare in the Middle East throughout the 1900s, and were believed to have been extirpated by the mid-1950s; however, recent sightings of tracks and a photo of a bear in western Syria, near the Lebanon border, suggests either a relict population, vagrants from Turkey, or released captive animals (Garshelis et al. 2015).
Presently, Brown Bears occupy approximately 5,000,000 km² of the northwestern portion of North America, 1,200,000 km² of Europe (excluding Russia), and much of northern Asia. Resident populations are known to exist in 45 range countries. The largest numbers exist in Russia, the U.S. (Alaska), and Canada. Many populations in Europe and the more southerly portions of Asia and North America are small and isolated (Servheen et al. 1999, Swenson et al. 2000, Kaczensky et al. 2013). Very small numbers of brown bears still remain in several Asian countries such as Iraq and Nepal (Gurung 2004, Ridings 2006, Aryal et al. 2012). In Europe, Andorra was reoccupied in 2003 from bears reintroduced into the French Pyrenees. A few wandering individuals are periodically crossing into Switzerland from a reintroduced population in northern Italy, and into Lithuania from Latvia and Belarus, but not enough as yet to be considered as occupied range in these countries.
Native:Afghanistan; Albania; Armenia (Armenia); Austria; Azerbaijan; Belarus; Bosnia and Herzegovina; Bulgaria; Canada; China; Croatia; Estonia; Finland; France; Georgia; Greece; India; Iran, Islamic Republic of; Iraq; Italy; Japan; Kazakhstan; Korea, Democratic People's Republic of; Kyrgyzstan; Latvia; Macedonia, the former Yugoslav Republic of; Mongolia; Montenegro; Nepal; Norway; Pakistan; Poland; Romania; Russian Federation; Serbia (Serbia); Slovakia; Slovenia; Spain; Sweden; Tajikistan; Turkey; Ukraine; United States (Georgia); Uzbekistan
Possibly extinct:Bhutan; Lebanon
Regionally extinct:Algeria; Belgium; Denmark; Egypt; Germany; Hungary; Ireland; Israel; Jordan; Liechtenstein; Lithuania; Luxembourg; Mexico; Moldova; Monaco; Morocco; Netherlands; Palestinian Territory, Occupied; Portugal; San Marino; Tunisia; United Kingdom
|Range Map:||Click here to open the map viewer and explore range.|
The total number of brown bears on earth is estimated to exceed 200,000. Reliable population estimates (derived mainly from mark-recapture or resight, and modifications thereof) exist for several areas in North America and Europe (Miller et al. 1997, Swenson et al. 2000, Bellemain et al. 2005, Mowat et al. 2005, Proctor et al. 2010, 2012, Mowat et al. 2013), but few areas in Asia. Russia has the largest number of brown bears, believed to exceed 100,000, while estimates in the U.S. (mainly in Alaska) are around 33,000, Canada 25,000, and Europe (excluding Russia) 15,400. Whereas the species is relatively abundant in more northern parts of its distribution, the southern portions of the range are highly fragmented, with many small populations. In North America, the southern fringe has isolated populations ranging in size from near 700 bears in and around Yellowstone National Park (Haroldson et al. 2013) to approximately 25 individuals in the Cabinet Mountains of Montana (Proctor et al. 2004, Kendall et al. 2016) and even less, likely <10 bears, in some southern areas of British Columbia.
|Current Population Trend:||Stable|
|Habitat and Ecology:|
Brown Bears occupy a great variety of habitats from dry Asian steppes to Arctic shrublands to temperate rain forests. Their range overlaps that of both the American Black Bear (U. americanus) and Asiatic Black Bear (U. thibetanus), and also slightly that of the Polar Bear (U. maritimus). Elevationally they range from sea level to 5,000 m (Sathyakumar 2006). They occupy a greater diversity of habitats than any other species of bear and also exploit a large variety of food items. In terms of diet, they fall between the mainly plant-dependent ursids and the carnivorous polar bear (Mattson 1998, Sacco and Van Valkenburgh 2004). In North America, Brown Bears (Grizzly Bears) are more carnivorous where ungulates (especially in Arctic areas) or spawning salmon (coastal areas) are abundant (Mowat and Heard 2006). The productivity and density of brown bears varies enormously, corresponding with the productivity of their habitats and availability of salmon. Coastal areas of North America and Eastern Russia, with concentrations of spawning salmon, have high densities (>100 bears per 1,000 km²) of Brown Bears (Miller et al. 1997, Seryodkin 2006) with high reproductive rates (Hilderbrand et al. 1999). Deciduous and mixed forests of the Dinaric and Carpathian mountain ranges of Eastern Europe also host high bear densities with high reproductive rates (Kusak and Huber 1998, Frković et al. 2001). More moderate densities of bears (20-50 bears per 1,000 km²) occur across the interior mountain ranges of North America (McLellan 1994, Schwartz et al. 2003, McLellan 2011), Europe, and Asia where they forage on a great variety of grasses, herbs, roots, berries, nuts, as well as animal matter such as insects, mammals, and fish if available. Moderate densities of bears are also found across portions of the boreal forests of North American, Asia and Scandinavia (Bellemain et al. 2005). Lower densities (5‒15 bears per 1,000 km²) are found in dry, desert-like areas, alpine and sub-alpine areas, as well as areas where habitat availability and numbers of bears have been reduced by high human and domestic livestock densities (Nawaz 2007); however, in most of these areas (e.g., northern India, western China) density estimates are not available.
|Use and Trade:||
Where Brown Bears exist in a large, contiguous population, they are sometimes hunted for sport or killed for control purposes, and rates may sometimes be unsustainable at least over the short term (Lamb et al. submitted, McLellan 2015, McLellan et al. submitted). Estimates of sustainable exploitation are hampered by the difficulty and expense of obtaining reliable estimates of population size, natural mortality, and reproductive rates. Most countries do not have the resources to develop and implement adequate monitoring programs and sustainable management plans for brown bears. Moreover, even with such plans in place, illegal or unreported kill may equal or exceed the legal and supposedly sustainable kill (McLellan et al. 1999, McLellan 2015). This is apparently occurring in the Russian Far East and China, where brown bears are poached for the commercial trade in gall bladders and paws (Seryodkin 2006, Servheen 2013, Burgess et al. 2014). In portions of Alaska, the management goal is to reduce bear populations to encourage moose and caribou populations to expand for the benefit of hunters (Miller et al. 2011).
Although, as a whole, this species is secure with relatively large numbers and an expansive range, many small, isolated populations are threatened due to their low numbers and frequent contact with humans. These small populations tend to be found in remnant wild areas surrounded by more extensive human development. As wide-ranging omnivores, Brown Bears are attracted to areas with available human-related foods; being large and somewhat aggressive, these bears may threaten life and property (often agricultural products) and may be killed as a consequence. Areas of high human use that attract bears may serve as significant mortality sinks (Nielsen et al. 2004, 2006). Additionally, bears living near humans may be killed inadvertently (e.g., vehicle or train collisions), poached for parts or products, or killed by people hunting for other species. Even small numbers of bears removed from small populations can have adverse effects on population growth (Wakkinen and Kasworm 2004); conversely, preventing just a few deaths may avert a population decline (Wiegand et al. 1998, Garshelis et al. 2005). Where Brown Bears exist in a large, contiguous population, they are sometimes hunted for sport or killed for control purposes, and rates may sometimes be unsustainable at least over the short term (Lamb et al. submitted, McLellan 2015, McLellan et al. submitted). Estimates of sustainable exploitation are hampered by the difficulty and expense of obtaining reliable estimates of population size, natural mortality, and reproductive rates. Most countries do not have the resources to develop and implement adequate monitoring programs and sustainable management plans for Brown Bears. Moreover, even with such plans in place, illegal or unreported kill may equal or exceed the legal and supposedly sustainable kill (McLellan et al. 1999, McLellan 2015). This is apparently occurring in the Russian Far East, where brown bears are poached for the commercial trade in gall bladders and paws (Seryodkin 2006). In portions of Alaska, the management goal is to reduce bear populations to encourage moose and caribou populations to expand for the benefit of hunters (Miller et al. 2011). In addition to direct removal of brown bears, many other human activities (such as agriculture, plantation forestry, highways, hydroelectric and wind power developments, and human settlements) eliminate, fragment, or erode the value of bear habitat (Proctor et al. 2005, Waller and Servheen 2005, Proctor et al. 2012). Habitat fragmentation is a serious threat that isolates population units with deleterious demographic and genetic impacts (Proctor et al. 2005, 2012). With increasing human populations, the value of brown bear habitat is being degraded in many areas (e.g., Can and Togan 2004, Nawaz 2007), while in other areas (i.e. Turkey), fewer people are now living in rural areas and conditions are improving for Brown Bears.
In Europe, four of ten populations are Critically Endangered. However, the current public interest, financial investment and management actions, seem to presently secure most populations at least for their short to midterm survival. Portions of populations in some countries are less secure due to lower local acceptance and correspondingly high human-caused mortality rates. Almost half of the populations are currently growing, but to guarantee long-term survival, all present and potential future threats must be kept in check. The key threats for bears in Europe were identified as: habitat loss due to infrastructure development, disturbance, low acceptance, poor management structures, intrinsic factors, accidental mortality and persecution. Most threats were expected to increase in the future (Kaczensky et al. 2013).
Conservation actions for Brown Bears vary greatly among nations and regions within nations. Large populations of this species (in Russia, Japan, Canada, Alaska, and parts of eastern and northern Europe) are legally hunted, and thus managed as a game animal. Hunting regulations, usually designed to ensure a sustainable harvest of bears, vary among areas but often involve a lottery for a limited number of permits (a quota system) and restricted season length. Most small populations are protected by national laws and international agreements, with varying degrees of enforcement. Brown Bear as a species is listed under CITES Appendix II; the populations in Bhutan (if they exist), China, and Mongolia, and those classified as the subspecies U. a .isabellinus (northern India, Pakistan, Afghanistan, to Kazakhstan and Gobi desert) are listed under Appendix I. Furthermore, international trade in bears and bear products from certain populations in countries of the European Union is restricted by EU regulations. In parts of the U.S., small populations of Grizzly Bears have increased under protection of the Endangered Species Act (U.S. Fish and Wildlife Service 2005). Reintroductions, population augmentations, and connectivity management also have helped to restore numbers and geographic range in several locations in the U.S., southern Canada, and Western Europe (Clark et al. 2002, Kasworm et al. 2007). There are numerous protected areas around the world with Brown Bears, but few are large enough to support a viable population; therefore, Brown Bear conservation must be integrated with many other human land-uses (Herrero 1994, Nielsen et al. 2006). Some countries have rules or management guidelines designed to reduce human impacts on Brown Bears and their habitat, whereas in other countries bear management protocols and regulations are limited or non-existent (Servheen et al. 1999, Zedrosser et al. 2001, Kaczensky et al. 2013).
In Europe most of the bear populations are protected by the Habitat Directive (Habitat Directive; http://eur-lex.europa.eu/legal-content/EN/TXT/HTML/?uri=CELEX:01992L0043-20070101&from=EN) which is compulsory for all EU countries. Portions of populations that fall within EU countries are not allowed to be classified as game animals. However, Sweden, Finland, Romania, Estonia, Bulgaria, Slovenia, Slovakia, Bulgaria and Croatia currently use derogations under article 16 of the Habitat Directive to allow a limited kill of bears by hunters. Bosnia and Herzegovina and Norway manage bears as a game species with annual quotas as they are only limited by the Bern Convention. Trophy hunting of bears enhances local acceptance of these animals by the public in some populations (Knott et al. 2013). Nearly all European countries have some form of bear management plan, action plan or bear management strategy. However, in a number of countries these documents have not been adequately implemented.
Abbas, F., Bhatti, Z.I., Haider, J., and Mian, A. 2015. Bears in Pakistan: distribution, population biology and human conflicts. Journal of Bioresource Management 2: 1-13.
Aryal, A., Sathyakumar, S., and Schwartz, C.C. 2010. Current status of brown bears in the Manasalu Conservation Area, Nepal. Ursus 21: 109-114.
Bellemain, E., Swenson, J. E., Tallmon, D., Taberlet, P. and Brunberg, S. 2005. Estimating population size of elusive animals with DNA from hunter-collected feces: Four methods for brown bears. Conservation Biology 19: 150-161.
Brown, D. E. 1985. The grizzly in the southwest. Documentary of an extinction. University of Oklahoma Press, Norman, Oklahoma, USA.
Burgess, E.A., Stoner, S.S., and Foley, K.E. 2014. Brought to bear” an analysis of seizures across Asia (2000-2001). TRAFFIC, Petaling Jaya, Selangor, Malayasia.
Can, O. E. and Togan, I. 2004. Status and management of brown bears in Turkey. Ursus 15(1): 48-53.
Chapron, G., Kaczensky, P., Linnell, J.D.C., von Arx, M., Huber, D., Andrén, H., Vicente López-Bao, J., Adamec, M., Álvares, F., Anders, O., Balčiauskas, L., Balys, V., Bedő, P., Bego, F., Blanco, J. C., Breitenmoser, U., Brøseth, H., Bufka, L., Bunikyte, R., Ciucci, P., Dutsov, A., Engleder, T., Fuxjäger, C., Groff, C., Holmala, K., Hoxha, B., Iliopoulos, Y., Ionescu, O., Jeremić, J., Jerina, K., Kluth, Knauer, F., Kojola, I., Kos, I., Krofel, M., Kubala, J., Kunovac, S., Kusak, J., Kutal, M., Liberg, O., Majić, A., Männil, P., Manz, R., Marboutin, E., Marucco, F., Melovski, D., Mersini, K., Mertzanis, Y., Mysłajek, R.W., Nowak, S., Odden, J., Ozolins, J., Palomero, G., Paunović, M., Persson, J., Potočnik, H., Quenette, P.-Y., Rauer, G., Reinhardt, I., Rigg, R., Ryser, A., Salvatori, V., Skrbinšek, T., Stojanov, A., Swenson, J.E., Szemethy, L., Trajçe, A., Tsingarska-Sedefcheva, E., Váňa, M., Veeroja, R., Wabakken, P., Wölfl, M., Wölfl, S., Zimmermann, F., Zlatanova, D., and Boitani, L. 2014. Recovery of Large carnivores in Europe's modern human-dominated landscapes. Science 346: 1517-1519.
Clark, J. D., Huber, D. and Servheen, C. 2002. Reintroducing bears: lessons and challenges. Ursus 13: 153-163.
Frković, A., Huber, D. and Kusak, J. 2001. Brown bear litter sizes in Croatia. Ursus 12: 103-106.
Gärdenfors, U., Hilton-Taylor, C., Mace, G., and Rodríguez, J.P. 2001. The application of IUCN Red List Criteria at Regional levels. Conservation Biology 15: 1206-1212.
Garshelis, D.L., Gibeau, M.L. and Herrero, S. 2005. Grizzly bear demographics in and around Banff National Park and Kananaskis Country, Alberta. Journal of Wildlife Management 69: 277-297.
Garshelis, D.L., Steinmetz, R., Hajjar, I., and Tabbaa, D. 2015. Brown bear(s) do exist in Syria: fuzzy photo generates much interest. International Bear News 24: 5-8.
Gong, J. and Harris, R.B. 2006. The status of bears in China. Understanding Asian bears to secure their future, pp. 50-56. Japan Bear Network, Ibaraki, Japan.
Guilday, J.E. 1968. Grizzly bears from eastern North America. The American Midland Naturalist 79: 247-250.
Gurung, M. K. 2004. Brown bear observation in Damodar Kunda Valley, Mustang District, Nepal. International Bear News 13(4): 12-14.
Hamdine, W., Thevenot, M. and Michaux, J. 1998. Recent history of the brown bear in the Maghreb. Comptes Rendus de l'Academie des Sciences - Serie III: Sciences de la Vie 321: 565-570.
Haroldson, M. A., F. T. van Manen, and D. D. Bjornlie. 2013. Estimating number of females with cubs-of-the-year. In: F. T. van Manen, M. A. Haroldson, and K. West (eds), Yellowstone grizzly bear investigations: annual report of the Interagency Grizzly Bear Study Team, 2012. U.S., pp. 11–18. Geological Survey, Bozeman, Montana, USA.
Herrero, S. 1994. The Canadian national parks and grizzly bear ecosystems: the need for interagency management. International Conference on Bear Research and Management 9: 7-21.
Hilderbrand, G. V., Jacoby, M. E., Schwartz, C. C., Arthur, S. M., Robbins, C. T., Hanley, T. A. and Servheen, C. 1999. The importance of meat, particularly salmon, to body size, population productivity, and conservation of North American brown bears. Canadian Journal of Zoology 77: 132-138.
IUCN. 2012. Guidelines for Application of IUCN Red List Criteria at Regional and National Levels: Version 4.0. IUCN, Gland, Switzerland and Cambridge, UK.
IUCN. 2016. The IUCN Red List of Threatened Species. Version 2016-3. Available at: www.iucnredlist.org. (Accessed: 07 December 2016).
Kaczensky P., G. Chapron, G., von Arx, M., Huber, D., Andrén, C.H., and Linnell, J. 2013. Status, management and distribution of large carnivores – bear, lynx, wolf and wolverine in Europe. Instituto di Ecologia Applicata and with contributions of the IUCN/SSC Large Carnivore Initiative for Europe.
Kasworm, W.F., Proctor, M.F., Servheen, C., and Paetkau, D. 2007. Success of grizzly bear population augmentation in northwest Montana. Journal of Wildlife Management 71: 1261–1266.
Kendall, K.C., Macleod, A.C., Boyd, K.L., Boulanger, J., Royle, J.A., Kasworm, W.F., Paetkau, D., Proctor, M.F., Annis, K., and Graves, T.A. 2016. Density, distribution, and genetic structure of grizzly bears in the Cabinet-Yaak Ecosystem. Journal of Wildlife Management 80: 314-331.
Knott. E.J., Bunnefeld, N.,Huber, D., Reljić, S., Kereži, V., and Milner-Gulland, E.J. 2013. The potential impacts of changes in bear hunting policy for hunting organisations in Croatia. European Journal of Wildllife Research.
Kusak, J. and Huber, D. 1998. Brown bear habitat quality in Gorski kotar, Croatia. Ursus 10: 281-291.
Lamb, C.T., Mowat, G., McLellan, B.M., Nielsen, S.E., and Boutin, S. Submitted. Forbidden fruit: Human settlement and abundant fruit create an ecological trap for an apex omnivore. Journal of Animal Ecology .
Loring, S., and Spiess, A. 2007. Further documentation supporting the former existence of grizzly bears (Ursus arctos) in Northern Quebec-Labrador. Arctic 60: 7-16.
Mace, R.D., Carney, D.W., Chilton-Radant, T., Courville, S.A., Haroldson, M.A., Harris, R.B., Jonkel, J., McLellan, B.N., Madel, M., Manley, R.L., Schwartz, C.C., Servheen, C., Stenhouse, G., Waller, J.S., and Wenum, E. 2012. Grizzly bear population vital rates and trend in the Northern Continental Divide Ecosystem, Montana. Journal of Wildlife Management 76: 119–128.
Manlius, N. 1998. L’Ours brun en Égypte [The brown bear of Egypt]. Écologie 29: 565-581.
Mano, T. 2006. The status of brown bears in Japan. Understanding Asian bears to secure their future, pp. 111-121. Japan Bear Network, Ibaraki, Japan.
Mattson, D. J. 1998. Diet and morphology of extant and recently extinct northern bears. Ursus 10: 479–96.
Mattson, D. J. and Merrill, T. 2002. Extirpations of grizzly bears in the contiguous United States, 1850-2000. Conservation Biology 16: 1123-1136.
McLellan, B. N. 1994. Density dependent population regulation of brown bears. International Conference on Bear Research and Management Monograph Series 3: 15-24.
McLellan, B.N. 2011. 2011. Implications of a high-energy and low-protein diet on body composition, fitness, and competitive abilities of black (Ursus americanus) and grizzly (Ursus arctos) bears. Canadian Journal of Zoology 89: 546-558.
McLellan, B.N. 2015. Some mechanisms underlying variation in vital rates of grizzly bears on a multiple use landscape. Journal of Wildlife Management 79: 749-765.
McLellan, B.N., Hovey, F.W., Mace, R.D., Woods, J.G., Carney, D.W., Gibeau, M.L., Wakkinen, W.L., and Kasworm, W.F. 1999. Rates and causes of grizzly bear mortality in the interior mountains of British Columbia, Alberta, Montana, Washington, and Idaho. Journal of Wildlife Management 63: 911-920.
McLellan, B.N., Mowat, G., Hamilton, A.N., and Hatter, I.W. Submitted. Is the grizzly bear hunt in British Columbia sustainable? Journal of Wildlife Management.
Miller, S.D., Schoen, J.W., Faro, J., and Klein, D.R. 2011. Trends in intensive management of Alaska’s grizzly bears, 1980-2010. Journal of Wildlife Management 75: 1243-1252.
Miller, S.D., White, G.C., Sellers, R.A., Reynolds, H.V., Schoen, J.W., Titus, K., Barnes, V.G., Smith, R.B., Nelson, R.R., Ballard, W.B. and Schwartz, C.C. 1997. Brown and black bear density estimation in Alaska using radiotelemetry and replicated mark-resight techniques. Wildlife Monographs 133: 5-55.
Mowat, G. and Heard, D. C. 2006. Major components of grizzly bear diet across North America. Canadian Journal of Zoology 84: 473-489.
Mowat G, Heard, D.C., and Schwarz, C.J. 2013. Predicting grizzly bear density in western North America. PLoS ONE 8(12).
Mowat, G., Heard, D. C., Seip, D. R., Poole, K. G., Stenhouse, G. and Paetkau, D. W. 2005. Grizzly Ursus arctos and black bear U. americanus densities in the interior mountains of North America. Wildlife Biology 11: 31-48.
Nawaz, M. A. 2007. Status of the brown bear in Pakistan. Ursus 18: 89–100.
Nawaz. M.A., Swenson, J., and Zakaria, V. 2008. Pragmatic management increases a flagship species the Himalayan brown bears, in Pakistan’s Deosai National Park. Biological Conservation 141: 2230-2241.
Nielsen, S. E., Herrero, S., Boyce, M. S., Mace, R. D., Benn, B., Gibeau, M. L. and Jevons, S. 2004. Modelling the spatial distribution of human-caused grizzly bear mortalities in the Central Rockies ecosystem of Canada. Biological Conservation 120: 101-113.
Nielsen, S. E., Stenhouse, G. B. and Boyce, M. S. 2006. A habitat-based framework for grizzly bear conservation in Alberta. Biological Conservation 130: 217-229.
Proctor, M. F., Mclellan B. N., Strobeck, C. and Barclay, R. M. R. 2005. Genetic analysis reveals demographic fragmentation of grizzly bears yielding vulnerably small populations. Proceedings of the Royal Society of London B Biological Sciences 272: 2409-2416.
Proctor, M.F., Paetkau, D., McLellan, B.N., Stenhouse, G.B., Kendall, K.C., Mace, R.D., Kasworm, W.F., Servheen, C., Lausen, C.L., Gibeau, M.L., Wakkinen, W.L., Haroldson, M.A., Mowat, G., Apps, C.D., Ciarniello, L.M., Barclay, R.M.R., Boyce, M.S., Strobeck, C., and Schwartz, C.C. 2012. Population fragmentation and inter-ecosystem movements of grizzly bears in western Canada and the northern United States. Wildlife Monographs 180: 1-46.
Proctor, M. F., Servheen, C., Miller, S. D., Kasworm, W. F. and Wakkinen, W. L. 2004. A comparative analysis of management options for grizzly bear conservation in the US-Canada trans-border area. Ursus 15: 145-160.
Ridings, C. 2006. Green bear in the desert. International Bear News 15(2): 12-13.
Sacco, T. and Van Valkenburgh, B. 2004. Ecomorphological indicators of feeding behaviour in the bears (Carnivora: Ursidae). Journal of Zoology (London) 263: 41-54.
Sathyakumar, S. 2006. The status of brown bears in India. Understanding Asian bears to secure their future, pp. 7-11. Japan Bear Network, Ibaraki, Japan.
Sathyakumar S., Kaul, R., Ashraf, N.V., Mookerjee, A., and Menon, V. 2012. National bear conservation and welfare action plan. Ministry of Environment and Forests, Wildlife Institute of India and Wildlife Trust of India.
Schwartz, C.C.,Haroldson, M.A., G.C., Cherry, S., Keating, K.A., Moody, D., and Servheen, C. 2006. Temporal, spatial, and environmental influences on the demographics of grizzly bears in the greater Yellowstone Ecosystem. Wildlife Monographs 161(1).
Schwartz, C. C., Miller, S. D. and Haroldson, M. A. 2003. Grizzly bear. In: G. A. Feldhamer, B. C. Thompson, and J. A. Chapman (eds), Wild mammals of North America: biology, management, and conservation, pp. 556-586. Johns Hopkins University Press, Baltimore, Maryland, USA.
Servheen, C. 2013. Bear paw smuggling into China continues. International Bear News 22(3): 24.
Servheen, C., Herrero, S. and Peyton, B. 1999. Bears. Status Survey and Conservation Action Plan. IUCN/SSC Bear and Polar Bear Specialist Groups, Gland, Switzerland.
Seryodkin, I. 2006. The biology and conservation status of brown bears in the Russian Far East. Understanding Asian bears to secure their future, pp. 79-85. Japan Bear Network, Ibaraki, Japan.
Swenson, J.E., Sandergren, F., and Söderberg, A. 1998. Geographic expansion of an increasing brown bear population: evidence for presaturation dispersal. Journal of Animal Ecology 67: 819–82.
Swenson, J., Gerstl, N., Dahle, B. and Zedrosser, A. 2000. Action plan for the conservation of the brown bear in Europem (Ursus arctos). Council of Europe, Strasbourg, France.
Tumendemberel, O., Proctor, M., Reynolds, H., Boulanger, J., Luvsamjamba, A., Tserenbataa, T., Batmunkh, M., Craighead, D., Yanjin, N., Paetkau, D. 2015. Gobi bear abundance and inter-oases movement survey, Gobi Desert, Mongolia. Ursus 26: 129-142.
U.S. Fish and Wildlife Service. 2005. Endangered and threatened wildlife and plants; Designating the greater Yellowstone ecosystem population of grizzly bears as a distinct population segment; removing the Yellowstone distinct population segment of grizzly bears from the federal list of endangered and threatened wildlife. United States Federal Register. 70: 69854-69884.
Wakkinen, W. L. and Kasworm, W. F. 2004. Demographics and population trends of grizzly bears in the Cabinet-Yaak and Selkirk ecosystems of British Columbia, Idaho, Montana, and Washington. Ursus 15: 65-75.
Waller, J. S. and Servheen, C. 2005. Effects of transportation infrastructure on grizzly bears in northwestern Montana. Journal of Wildlife Management 69: 985-1000.
Wiegand, T., Stephan, T., Naves, J. and Fernandez, A. 1998. Assessing the risk of extinction for the brown bear (Ursus arctos) in the Cordillera Cantabrica, Spain. Ecological Monographs 68: 539-570.
Zedrosser, A., Dahle, B., Swenson, J. E. and Gerstl, N. 2001. Status and managment of the brown bear in Europe. Ursus 12: 9-20.
|Citation:||McLellan, B.N., Proctor, M.F., Huber, D. & Michel, S. 2016. Ursus arctos. The IUCN Red List of Threatened Species 2016: e.T41688A45034772.Downloaded on 30 April 2017.|
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