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Capra aegagrus

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Taxonomy [top]

Kingdom Phylum Class Order Family
ANIMALIA CHORDATA MAMMALIA CETARTIODACTYLA BOVIDAE

Scientific Name: Capra aegagrus
Species Authority: Erxleben, 1777
Common Name(s):
English Bezoar, Wild Goat
French Chèvre Égagre
Taxonomic Notes: The Domestic Goat and the Wild Goat are treated here as separate species (following inter alia Shackleton 1997), named Capra hircus and Capra aegagrus respectively. These taxa are sometimes considered to be conspecific, in which case the name Capra aegagrus has generally been used to refer to the wild species and its domesticated form, although some authors use the name Capra hircus for both the wild species and its domestic descendants (see Gentry et al. 1996, BZN 2003, Gentry et al. 2004 and Wilson and Reeder 2005).

"Wild goats" and "wild sheep" found on Mediterranean islands are generally recognized to have been introduced by humans (Shackleton 1997, Wilson and Reeder 2005), and genetic and archaeozoological studies suggest that they are feral populations of ancient domestic stocks (e.g. ,Groves 1989, Vigne 1994, Hiendleder et al. 1998, Manceau et al. 1999, Kahila bar-Gal et al. 2002). Consequently, such taxa should be included in the respective domestic species (Capra hircus, Ovis aries) and not as subspecies of the wild taxa (as proposed by Gentry et al. 1996, Gentry et al. 2004, and Gippoliti and Amori 2004).

Assessment Information [top]

Red List Category & Criteria: Vulnerable A2cd ver 3.1
Year Published: 2008
Date Assessed: 2008-06-30
Assessor(s): Weinberg, P., Jdeidi, T., Masseti, M., Nader, I., de Smet, K. & Cuzin, F.
Reviewer(s): Festa-Bianchet, M. & Harris, R. (Caprinae Red List Authority)
Justification:
Listed as Vulnerable because of a population decline, estimated to be more than 30% over the last three generations, inferred from over-exploitation, shrinkage in distribution, and habitat destruction and degradation.
History:
1996 Vulnerable

Geographic Range [top]

Range Description: The wild goat ranges discontinuously from central Afghanistan and southern Pakistan, west through Iran, western Turkmenistan, northern Iraq, the Caucasus region (Armenia, Azerbaijan, northeastern Georgia, and southern Russia), as far as southwestern Turkey. It once occurred in Jordan, Lebanon and Syria, but is now extinct in these countries (Grubb, 2005). It also occurred in Israel before 10,000 ago (Dayan et al. 1986).

Its range in particular countries and regions is as follows:

Afghanistan
It is probably confined to the Hazarajat and Uruzgan mountains in central Afghanistan, including the arid Feroz Koh and Siyah Koh in the headwaters of the Hari Rud, Farah Rud, Helmand and Arghandab rivers. However, no animals were observed by FAO or WWF survey teams in the 1970s, but horns and skulls were occasionally seen at shrines and grave sites. One captive animal seen in 1975 in a private zoo in Kandahar, was reputedly caught in the nearby mountains. The species had probably been reduced to a small portion of its former range by the late 1970s (Habibi 1977). Its current status in the country is unknown.

Caucasus
One subspecies of the wild goat is recognised in the Caucasus. The distribution of C. a. aegagrus is in two separate parts. It occurs in forested areas along northern slopes of the Greater Caucasus mountains from the Upper Argun river, in Chechen-Ingushetia and in Georgia, up to the headwaters of the Jurmut river (around 42°30'N, 46°E) in Daghestan, with an isolated population on the southern slopes of Mount Babadagh, Azerbaijan. It also occurs in the drier, more open habitats in the Caucasus Minor mountains in both Azerbaijan and Armenia, south and east of Sevang lake, namely the Shakhdagh, Mrovdagh, Karabakh, Gegam, Vardenis and Zangezur ranges, and on the Delidagh massif between 39° to 40°30'N and 45° to 46°39'E.

Iran
The wild goat is widely distributed throughout Iran wherever large areas of rocky terrain are available. This includes not only mountainous areas, but also cliffs along the seashore, in deciduous forested areas of the north, and in areas of the central desert.

Iraq
If it still occurs in Iraq, it would most likely be found in the Zagros mountains in the extreme north and along the northeastern border with Iran. Nothing is known of current distributions.

Lebanon
The wild goat used to be relatively common in Barouk, the Ammiq mountains and on Mount Harmon, northern Lebanon. However, by the early 1900s it was extinct in Lebanon (Kumerloeve 1975; Harrison 1968).

Pakistan
The present range of Sind wild goat (C. a. blythi) is the Baluchistan plateau and its foothills in south-western Pakistan. Populations are scattered on arid mountain ranges that are isolated by lowlands of southern Baluchistan and Sind. The range includes the low Mekran coastal range (District Gwadar), areas up to 3,250 m asl in the Koh-i-Maran range south of Quetta (District Kalat), and also the Kirthar range (Districts of Dadu and Las Bela) (Roberts 1977, Schaller 1977, Bollmann 1989). A second subspecies, the Chiltan goat (C. a. chialtanensis) was restricted in the early 1970s to four or five populations in the accessible mountain ranges (Chiltan, Murdar, Koh-i-Maran and Koh-i-Gishk ranges) south of Quetta (Roberts 1977, Schaller 1977). Today, these have been reduced, principally by uncontrolled hunting, to only one surviving population in the Hazarganji-Chiltan National Park (Districts of Quetta and Kalat).

Syria
Wild goat was reported in northern Syria, in the mountains north of Dimasq, and must once have occurred in the western mountains as well. However, it is now believed to be extinct (Harrison 1968).

Turkey
The wild goat ranges widely in Turkey, east from the Datca peninsula, through the Taurus and Anti-Taurus mountains in the mountainous regions of southeastern, eastern and northeastern Anatolia (Kence 1987).

Turkmenistan
The Turkmen wild goat (C. a. blythi [= turcmenica]), occurs in scattered populations in the central Kopet Dagh along the border between Turkmenistan and Iran between about 37°20'N, 55°20'E and 38°N, 57°20'E, and in the Large Balkhan (Bolshye) north of Nebit Dagh about 39°30'N, 50°30'E. It is not known whether this subspecies still inhabits the Small Balkhan (Malye).
Countries:
Native:
Armenia (Armenia); Azerbaijan; Georgia; Iran, Islamic Republic of; Pakistan; Russian Federation; Turkey; Turkmenistan
Regionally extinct:
Jordan; Lebanon; Syrian Arab Republic
Range Map: Click here to open the map viewer and explore range.

Population [top]

Population: The global population of wild goat has not been estimated. Although the species ranges very widely, it is probably extremely rare or absent in much of its mapped range. In some places it is clearly decreasing rapidly. However, there is also evidence of localised population recovery when adequate protection is in place. The population trend across its range is likely to be a significant decrease, perhaps as much as 30% over three generations. There is an urgent need for updated information on the status of this species. Specific information on country-level abundance is as follows:

Afghanistan
No estimate of numbers available, but the species is probably now very rare in this country.

Caucasus
It inhabits mainly forested habitat, so accurate censuses are difficult to make. The total population estimate for wild goat was between 3,500 and 4,000 individuals in the late 1980s, with 1,500 in the Greater Caucasus (Arabuli 1989; Bathiyev 1989; Prilutskaya and Pishvanov 1989) and the rest in the Caucasus Minor, where more than half (1,000 to 1,250) lived in the southern half of the Zangezur range (Kuliyev 1981). At the end of 1990s, 2,500 animals were estimated for Daghestan alone (Nasrulayev, 2003), but numbers were declining rapidly, by 50% in three years (Magomedov, Omarov, Nasrulayev, 2001). The overall population trend for this subspecies is negative, and in recent years the rate of decline has increased.

Iran
No estimates of total numbers are currently available. However, 1991 estimates are available for Golestan National Park - 2,500 animals, and for Alborz-Markazy Protected Area - 4,000 animals.

Iraq
There are no population estimates, but the species is probably extremely rare, if it survives at all.

Pakistan
For C. a. blythi there is no overall population estimate. Most survive in small inaccessible areas in isolated populations (Virk 1991). However, reasonable numbers are reported for the Dhrun and Hingol areas (Ahmad and Beg 1989). Kirthar National Park probably contains the largest population of Sind wild goat in the country. In the Karchat Mountains, which are withinf Kirthar National Park, the population has increased from between 400 and 500 when the establishment park was established in 1973 (Schaller and Laurie 1974) to around 950 to 1,050 (Bollmann 1989). For the whole Park, Mirza and Asghar (1980) estimated a total of 1,480 goats, while Kermani and Khan (1985) gave an optimistic number of 4,000 animals. According to Bollmann's (unpubl. data) observations in 1987, the total population inhabiting the Park was between 1,500 and 2,000 goats. The adjacent Surjan-Sumbak-Eri-Hothiano Game Reserve contained another 900 to 1,100 (Mirza and Asghar 1980), resulting in a total estimate of 2,400 to 3,100 wild goat for Sind Province. Depending on the study area, densities ranged between 0.5 to 0.65 animals/km² (over 3,087 km² in Kirthar National Park; Bollmann, unpubl. data) and 11.8 to 16.3 animals/km² (over 60 km² in Karchat Mountains; Edge and Olsen-Edge 1990). However, the overall density within the species' distribution area in Pakistan is probably well below 0.5 wild goat/km². For C. a. chialtanensis the single population totalled ca. 168 animals in 1975 (Mirza 1975). Due to rigid protection following establishment of the National Park in 1980, the population had increased to 480 animals by 1990 (Baluchistan Forest Dept. Records, cited in Virk 1991) but this improvement may not have continued. There is an urgent need to updated information on the abundance of the wild goat in Pakistan.

Turkey
It is declining in Turkey throughout its range, and the total population is believed to be less than 10,000 mature individuals, with no subpopulation larger than 1,000 mature individuals.

Turkmenistan
Korshunov (1986), whose data are the most reliable, estimated that the total population was up to 7,000 animals.
Population Trend: Decreasing

Habitat and Ecology [top]

Habitat and Ecology: The wild goat inhabits mountainous areas, where there is a mixture of rocky outcrops (including scree slopes) and vegetation (shrubby thickets (maquis) or conifer forests). It tends to be found in relatively arid habitats, though it is a forest species in the Caucasus. It is herbivorous, feeding on grasses, herbaceous plants and shrubs.
Systems: Terrestrial

Use and Trade [top]

Use and Trade: This species is hunted mostly for food and sport, though the hides and other body parts may also be used.

Threats [top]

Major Threat(s): The major threats to this species are poaching, competition for food with domestic livestock, and disturbance and habitat loss from logging and land clearing.

Afghanistan
Overhunting and colonisation of their habitat by livestock resulted in depleted numbers prior to 1979, with small bands of wild goats forced into the most inaccessible parts of the mountain ranges.

Caucasus
Main threats in the Greater Caucasus are poaching and logging, since animals are mostly confined to forested areas. In the Caucasus Minor, there is also competition with livestock. The state of war between Azerbaijan and Armenia has a negative impact upon the main local populations of the wild goat.

Iran
Poaching, competition for food with domestic livestock, and disturbance and habitat loss from logging and land clearing, are major threats.

Lebanon
The extinction of wild goat in Lebanon was caused by large scale habitat destruction and the disregard of hunting regulations.

Pakistan
Within its range, the species is only locally abundant and under successful protection only in a few areas (e.g., in Sind Province). Most of the animals are in scattered populations on mountain ranges isolated from each other by lowlands. Consequently, they are at risk, especially because local people and nomadic tribes graze their domestic stock on most of the mountain ranges used by wild goats, and because hunting is still widespread.

Syria
Extinction was probably caused by habitat destruction and hunting.

Turkey
It is declining in Turkey principally due to over-hunting.

Conservation Actions [top]

Conservation Actions: Afghanistan
Reconnaissance surveys were conducted to evaluate the species? distribution patterns in the mid 1970s. The work was discontinued after the start of the civil war. Surveys to determine the current status and distribution of the species in the country and to develop conservation strategies.

Former Soviet Union
C. a. aegagrus is protected by law, and was included in Category II in the Red Data Book of the USSR (Borodin 1984), Georgia (1982) and Russia (1983). No more than 200 occur in at least three protected areas. The wild goat is found in Tushetian Reserve in the Greater Caucasus, and in the Caucasus Minor, in the Geigel (Azerbaijan) and the Khosrov (Armenia) Nature Reserves (Sokolov and Syroyechkovsky 1990). It also occurs in Ordubad Sanctuary (Nakhichevan), and periodically in ShikakhokhNature Reserve (Armenia). Turkmen wild goat (C. a. blythi [= turcmenica]) is also protected. It was included in Class II in the USSR Red Data Book (Borodin 1984). Kopet Dagh Nature Reserve is the only protected area with this subspecies. The Turkmenistan Government planned to allow hunts for 20 goats in 1993-94; the current licensed hunting situation is not clear. Throughout its range in the former Societ Union, more strict and effective measures are needed to control of poaching. Most local people in the Caucasus consider wild goats and other species as a source of meat, hides and horns, so stricter prohibition measures alone will not be sufficient. Public awareness programmes are also needed. It is essential to protect the forest habitat of this in Daghestan and elsewhere in the Caucaus. An Ordubad National Park has been established on the southern flank of Zangezur Ridge in Nakhichevan (Caucasus Minor) in an important site for the species. At least two new protected areas are needed, one in western Daghestan, and the other in the Large Balkhan. In Turkmenistan, the wild goat should be re-introduced into the Siunt-Khasardag Nature Reserve in the western Kopet Dagh, and into the Badkhyz Nature Reserve in the Gyaz-Gedyk range.

Iran
The wild goat occurs in several protected areas where hunting is prohibited and livestock grazing is strictly controlled. It is found in seven National Parks, 11 Wildlife Refuges, and 34 Protected Areas throughout the country. They can be hunted under licence outside protected areas between September and February each year, but more strict enforcement of hunting regulations is needed.

Iraq
Surveys are needed to determine the status and distribution of this species, and in particular whether or not any populations remain in the country.

Lebanon
A captive breeding plan and subsequent re-introduction program has been proposed, in conjunction with one in Syria.

Pakistan
In addition to the Kirthar and Hazarganji-Chiltan National Parks, there are several reserves providing different levels of protection to wild goats in the country, but for many, no management plans have been developed and most lack even supervisory staff (Virk 1991). Protected areas with Sind wild goat include: Baluchistan - Las Bela District: Hingol National Park, Dureji Wildlife Sanctuary, Khurkhera Wildlife Sanctuary (Baluchistan Forest Department 1990); Khuzdar District: Dhrun National Park (Baluchistan Forest Department 1990); Chorani Wildlife Sanctuary (Zoological Survey Department, no date); Gwadar District: Buzi Makola Wildlife Sanctuary (Zoological Survey Department, no date); Kharan District: Ragai Rakshan Wildlife Sanctuary, Ras Koh Wildlife Sanctuary (Baluchistan Forest Department 1990); Ras Koh Wildlife Sanctuary (Zoological Survey Department, no date); Chagae District: Gut Wildlife Sanctuary (Baluchistan Forest Department 1990). Kalat District: Koh-Egish Wildlife Sanctuary (Zoological Survey Department, no date); Dadu District: Kirthar National Park, Sumbak-Surjan-Eri-Hothiano Game Reserve (Mirza and Asghar 1980). A wildlife management plan for Kirthar National Park (Holloway and Khan 1973) recommended: 1) no extension of irrigated cultivation, 2) reduction of livestock grazing by the locals and immigrants, and 3) prohibition of tree felling, cutting branches for livestock fodder, poaching, stone quarrying and honey collection within the park. This plan is currently being revised. The world?s last remaining population of Chiltan goat is restricted to Hazarganji-Chiltan National Park (Quetta District; Zoological Survey Department, no date) where ca. 480 animals lived in 1990 (Forest Service reports cited in Virk 1991). Previous efforts to protect this Park included more administrative staff, control of livestock grazing, wood cutting, poaching, and construction of ca. 30 km boundary fence. This resulted in a marked recovery of the vegetation and an increase in the Chiltan goat population prior to 1990 (Baluchistan Forest Department Records, cited in Virk 1991). However, there are reports that conditions have recently deteriorated in this National Park. Conservation measures proposed for the species as a whole in Pakistan are as follows: 1) Implement the management ecommendations for Kirthar National Park. 2) Consider alternative livelihoods for the people who would be displaced by implementation Kirthar National Park?s management plan. 3) Develop a range management program for the Park based upon a system of zones of varying use-intensity by humans and livestock. 4) Implementation of such restrictions on the human population, must be accompanied not only by some form of compensation, but also include responsible involvement of the local community in conservation and agricultural programs. Virk (1991) emphasised the importance of participation of the local communities in the wildlife conservation process and gave a list of objectives for a local participatory approach, as well as a list of management goals. 5) Determine the current distribution and status of this species, and assess the remaining habitat before launching any further wild goat conservation projects in the Province of Baluchistan. 6) Consider implementing Virk?s (1991) three proposals for Hazarganji-Chiltan National Park: a) Enlarge the area of Hazarganji-Chiltan National Park by including the adjacent Kerkhasa State Forest (Ahmad 1988, cited in Virk 1991) to support any increase in the Chiltan goat population. b) Reintroduce and manage at least one herd of Chiltan goat in a formerly inhabited mountain range (e.g. Murdar, Koh-i-Maran). Such re-introductions would provide ways of presenting wildlife conservation to the public, and involve the local people in the conservation program. Natural dispersal from Hazarganji-Chiltan National Park to neighbouring mountain areas can be expected only over the long term and even then only with a high population density within the Park. c) Stop allowing hunters to take animals from the last population of Chiltan goat. Only when viable populations have been built up outside the National Park, could a trophy hunting program be considered as a possible means of generating funds for further conservation efforts. If conditions have deteriorated and the Park?s effectiveness has declined since 1990, obviously steps should be taken immediately to deal with the problems.

Syria
The proposed joint re-introduction project with Lebanon should be implemented.

Turkey
Much stricter conservation measures are needed to control hunting, and especially to secure the future of the larger populations.

Bibliography [top]

Ahmad, A. and Beg, A. R. 1989. A report on the floral and faunal characteristics of the newly established Hingol and Drun National Parks in Baluchistan. Wildlife Management Branch, Pakistan Forest Institute, Peshawar, Pakistan.

Arabuli, A. B. 1989. Contemporary status of rare ungulate species of Georgia. Ecology, Morphology, Use and Protection of Wild Ungulates, Part 1, pp. 28-29. Moscow, Russia.

Bathiyev, A. M. 1989. The wild goat in montane Chechen-Ingush. Nature and Economy of Chechen-Ingush ASSR, pp. 103-109. Chechen-Ingush State Publishers, Grozny.

Bollmann, K. 1989. Bestand, Populationsstruktur, Standortwahl und Aktivitaetsmuster der Bezoarziege (Capra aegagrus blythi) in Kirthar National Park, Sind, Pakistan. Diplomarbeit Universitaet Zurich.

Borodin, A. M. (ed.). 1984. USSR Red Data Book. Lesnaya Promyshlennost, Moscow, Russia.

Bulletin of Zoological Nomenclature. 2003. International Commission on Zoological Nomenclature, Opinion 2027 (Case 3010). Usage of 17 specific names based on wild species which are predated or contemporary with those based on domestic animals (Lepidoptera, Osteichthyes, Mammalia): conserved. Bulletin of Zoological Nomenclature 60: 81-84.

Dayan, T., Tchernov, E., Bar-Yosef, O. and Yom-Tov, Y. 1985. Animal exploitation in Ujrat El-Mehed, a Neolitic site in southern Sinai. Paleorient 12: 105-116.

Edge, W. D. and Olson-Edge, S. L. 1987. Ecology of wild goats and urial in Kirthar National Park. Final report. Montana Coop. Wildl. Res. Unit, University of Montana, Missoula, Montana, USA.

Gentry, A., Clutton-Brock, J. and Groves, C. P. 1996. Case 3010. Proposed conservation of usage of 15 mammal specific names based on wild species which are antedated by or contemporary with those based on domestic animals. Bulletin of Zoological Nomenclature 53: 28-37.

Gentry, A., Clutton-Brock, J. and Groves, C. P. 2004. The naming of wild animal species and their domestic derivatives. Journal of Archaeological Science 31: 645-651.

Gippoliti, S. and Amori, G. 2004. Mediterranean island mammals: are they a priority for biodiversity conservation? Biogeographia 25: 135?144.

Groves, C. P. 1989. Feral mammals of the Mediterranean islands: documents of early domestication. In: J. Clutton-Brock (ed.), The Walking Larder, pp. 46-58. Unwin-Hyman, London, UK.

Grubb, P. 2005. Artiodactyla. In: D. E. Wilson and D. M. Reeder (eds), Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), pp. 637-722. Johns Hopkins University Press, Baltimore, USA.

Habibi, K. 1977. The mammals of Afghanistan - their distribution and status. United Nations Development Programme - Food and Agriculture Organization of the United Nations - Dept. of Forest and Range, Ministry of Agriculture, Rome, Italy.

Harrison, D. L. 1968. The Mammals of Arabia. Ernest Benn Limited, London, UK.

Hiendleder, S., Mainz, K., Plante, Y. and Lewalski, H. 1998. Analysis of mitochondrial DNA indicates that domestic sheep are derived from two different ancestral maternal sources: no evidence for contributions from urial and argali sheep. Journal of Heredity 89: 113?120.

Holloway, C. M. and Khan, K. M. 1973. Management Plan for Kirthar National Park, Sind, Pakistan. Sind Wildlife Management Board and IUCN, Karachi, Pakistan and Gland, Switzerland.

Husband, T. P and Davis, P. B. 1984. Ecology and behaviour of the Cretan agrimi. Canadian Journal of Zoology 62: 411-420.

IUCN. 2008. 2008 IUCN Red List of Threatened Species. Available at: http://www.iucnredlist.org. (Accessed: 5 October 2008).

Kahila Bar-Gal, G., Smith, P., Tchernov, E., Greenblatt, C., Ducos, P., Gardeisen, A. and Horwitz, L. K. 2002. Genetic evidence of the origin of the agrimi goat (Capra aegagrus cretica). Journal of Zoology (London) 256: 369?377.

Kence, A. 1987. Biological Diversity in Turkey. Environmental Problems Foundation of Turkey, Ankara, Turkey.

Kermani, W. A. and Khan, K. M. 1985. Protected areas and local populations in Kirthar National Park, Pakistan. In: J. A. McNeely, J. W. Thorsell and S. R. Chalise (eds), People and Protected Areas in the Hindu Kush - Himalaya, pp. 59-61. King Mahendra Trust for Nature Conservation and International Centre for Integrated Mountain Development, Kathmandu, Nepal.

Korshunov, V. M. 1986. Numbers and biotopical distribution of Turkmen wild sheep and Turkmen wild goat in central Kopet Dag. Proceedings of the IV USSR Theriological Society 1: 249-250.

Kuliyev, S. M. 1981. The wild goat and the Daghestan rut in Azerbaijan [in Russian]. Candidate Thesis, University of Moscow.

Kumerloeve, H. 1975. Die saugetiere (Mammalia) der Turkei. Veroffentlichungen der Zoologischen Staatssammlung Munchen 18: 69-158.

Magomedov, M. R. D., Omarov, K. Z. and Nasrulaev, N. I. 2001. Specific character of anthropogenic impact upon mountain mammals of the East Caucasus. Sustainable development of mountain territories: problems of regional cooperation and regional policy of mountain areas: 379-381. Vladikavkaz, Russia.

Manceau, V., Després, L., Bouvet, J. and Taberlet, P. 1999. Systematics of the genus Capra inferred from Mitochondrial DNA sequence data. Molecular Phylogenetics and Evolution 13: 504?510.

Mirza, Z. B. 1975. A census of Chiltan markhor, Capra hircus in Chiltan Range, Quetta. Pakistan Journal of Zoology 7: 214-216.

Mirza, Z. B. and Asghar, M. 1980. Census of Sind ibex (Capra hircus blythi) and Gad (Ovis orientalis blanfordi) and some estimate of population of Chinkara (Gazella gazella) in Kirthar National Park and Sumbak Game Reserve, Sind. Pakistan Journal of Zoology 12: 268-271.

Nasrulaev, N. I. 2003. Regularities of structure-functional organization and peculiarities of ecology of wild goat (Capra aegagrus caucasica) population in Daghestan. Makhachkala.

Prilutskaya, L. I. and Pishvanov, Y. V. 1989. Range and numbers of of wild goat in Daghestan. Ecology, Morphology, Use and Protection of Wild Ungulates, Part 2, pp. 243-245. Moscow.

Roberts, T.J. 1977. The Mammals of Pakistan. Ernest Benn, London, UK.

Schaller, G.B. 1977. Mountain Monarchs: wild sheep and goats of the Himalaya. University of Chicago Press, Chicago, USA.

Schaller, G. B. and Laurie, A. 1974. Courtship behaviour of the wild goat. Zeitschrift für Säugetierkunde 39: 115-127.

Shackleton, D.M. 1997. Wild Sheep and Goats and Their Relatives: Status Survey and Conservation Action Plan for Caprinae. In: D. M. Shackleton (ed.), Wild sheep and goats and their relatives. Status survey and conservation action plan for Caprinae, IUCN/SSC Caprinae Specialist Group, Gland, Switzerland and Cambridge, UK.

Sokolov, V. E. and Syroyechkovsky, E. E. 1990. The Reserves of the Caucasus. Mysl, Moscow.

Vigne, J. D. 1994. Les transferts anciens de mammifères en Europe occidentale: histoires, mécanismes et implications dans les sciences de l'homme et les sciences de la vie. Colloques d'Histoire des Sciences zoologiques 5: 15-37.

Virk, A. T. 1991. Management plan for wild ungulates in Baluchistan, Pakistan. M. S. Thesis, University of Montana.

Wilson, D. E. and Reeder, D. M. 1993. Mammal Species of the World. A Taxonomic and Geographic Reference. Smithsonian Institution Press, Washington, DC, USA.

Wilson, D.E. and Reeder, D.M. 2005. Mammal Species of the World. Johns Hopkins University Press, Baltimore, MD, USA.

Zoological Survey Department, Govt. of Pakistan, in collaboration with Pakistan Agricultural Research Council, Islamabad and Pakistan Forest Institute, Peshawar. 1985. Wildlife of Pakistan Map 1: 1,950,OOO. Zoological Survey Departmen, Karachi.


Citation: Weinberg, P., Jdeidi, T., Masseti, M., Nader, I., de Smet, K. & Cuzin, F. 2008. Capra aegagrus. The IUCN Red List of Threatened Species. Version 2014.2. <www.iucnredlist.org>. Downloaded on 29 July 2014.
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