|Scientific Name:||Bos gaurus|
|Species Authority:||C.H. Smith, 1827|
Bos asseel Horsfield, 1851
Bos cavifrons Hodgson, 1837
Bos gaur Sundevall, 1846
Bos gaurus Lydekker, 1907 subspecies hubbacki
Bos gour Hardwicke, 1827
Bos subhemachalus Hodgson, 1837
Bubalibos annamiticus Heude, 1901
Gauribos brachyrhinus Heude, 1901
Gauribos laosiensis Heude, 1901
Gauribos mekongensis Heude, 1901
Gauribos sylvanus Heude, 1901
Uribos platyceros Heude, 1901
|Taxonomic Notes:||The International Commission on Zoological Nomenclature (2003) ruled that the name for this wild species is not invalid by virtue of being antedated by the name based on the domestic form. Therefore, IUCN considers the wild species of Gaur under Bos gaurus, while referring to the domestic form (Mythun, Mithan or Gayal) as Bos frontalis Lambert, 1804 (see Gentry et al. 2004).
Traditionally, three subspecies of Gaur have been recognized: Bos gaurus gaurus in India, Nepal, and Bhutan; B. g. readei in Myanmar (Burma), southern China, Lao PDR, Viet Nam, Cambodia, and Thailand north of the Isthmus of Kra (Lydekker 1903); and B. g. hubbacki in Thailand south of the Isthmus of Kra and in West Malaysia (Lydekker 1907). (In addition, Hubback (1937) thought that there may be two subspecies of Gaur in Malaysia, one with a well developed dewlap and one without.) However, this trifid arrangement, based largely on differences in coloration and size, is no longer widely recognized (Grzimek 1990, Corbet and Hill 1992).
Groves and Grubb (cited in National Research Council 1983) concluded that there were only two subspecies: Bos gaurus gaurus in India and Nepal and Bos gaurus laosiensis in Myanmar (Burma), Lao PDR, Viet Nam, Cambodia, Thailand, and West Malaysia (and presumably southern China). More recent work by Groves and colleagues (e.g., Groves 2003) also supports this division into two subspecies. Analysis of skull and horn measurements revealed little multivariate overlap between Indian and Southeast Asian specimens: Southeast Asian specimens are much bigger, with relatively shorter nasal bones, a less wide horn span and a narrower occiput; and in South-east Asian specimens the ascending branch of the premaxilla generally does not reach the nasal, whereas in Indian specimens it usually does. Specimens from Bhutan, Chittagong (Bangladesh), Upper Chindwin (north Myanmar), and Mogok (north Myanmar) were intermediate, but tended more towards the South-east Asian type. The locations of these specimens suggest that the Gaur in north-east India are also likely to be intermediate but more similar to southeast Asian animals than to the Gaur in the rest of India (C. Groves pers. comm. 1995). In recognition of these phenotypic differences, B. g. gaurus and B. g. laosiensis are provisionally accepted here pending further morphometric and genetic study.
The extinct subspecies B. g. sinhaleyus survived on Sri Lanka into historic times (Grubb 2005).
|Red List Category & Criteria:||Vulnerable A2cd+3cd+4cd ver 3.1|
|Assessor/s:||Duckworth, J.W., Steinmetz, R., Timmins, R.J., Pattanavibool, A., Than Zaw, Do Tuoc & Hedges, S.|
|Reviewer/s:||Burton, J. & Hedges, S. (Asian Wild Cattle Red List Authority)|
This species qualifies as Vulnerable under Criteria A2cd+3cd+4cd. The population decline in parts of the species' range especially Indochina and Malaysia, perhaps also Myanmar and China, is likely to be well over 70% over the last three generations (generation length estimated at 8–10 years), whereas in India and Nepal the overall decline rate is considerably lower. This reflects that in the various reasonably well-protected areas, population trends are stable, and in a few areas which have been neglected but are the focus of improved protection, populations are rebuilding. Weighting these by population size gives an overall decline of at least 30% over the last three generations. Similar reductions are projected, largely because the causative factors (especially wild meat hunting and hunting for the trade in horns in Southeast Asia) are still operative. Although the proportion of the global population which is in relatively well protected in India is increasing, and thus the averaged global decline rate ought to decrease, an ongoing decline of 30% (in the next three generations) or more is adjudged likely. The recent history in Malaysia proves that directed Gaur poaching can induce rapid declines in regions previously judged relatively safe. Poaching in Malaysia is undertaken by international gangs and it is likely that once returns in Malaysia drop they will move to other countries; the patterns in other high-value species in more advanced declines (e.g. southeast Asian otters) indicate that incursion even to southern India is likely. The species could potentially qualify for Vulnerable under Criteria C, although there are not enough quantitative data available
|Range Description:||Gaur historically occurred throughout mainland south and southeast Asia and Sri Lanka. It currently occurs in scattered areas in the following range states: Bhutan, Cambodia, China, India, Lao PDR, Malaysia (Peninsular Malaysia only), Myanmar, Nepal, Thailand, and Viet Nam, but is extinct in Sri Lanka and also, as a resident, apparently in Bangladesh (Grubb 2005; Md Anwarul Islam in litt. 2008; Hedges in prep.). The species is now seriously fragmented within its range, and the mapped distribution is generalised, especially in India, Myanmar, China and Malaysia.
The domesticated form of Gaur, considered by IUCN a separate species (Bos frontalis; Mythun, Mithan or Gayal), occurs in parts of India, China, and Myanmar as feral, semi-feral, and domestic animals. This animal is excluded from the red-listing considerations for Gaur.
Native:Bangladesh; Bhutan; Cambodia; China; India; Lao People's Democratic Republic; Malaysia (Peninsular Malaysia); Myanmar; Nepal; Thailand; Viet Nam
Regionally extinct:Sri Lanka
|Range Map:||Click here to open the map viewer and explore range.|
The global population is estimated to lie within 13,000–30,000 animals. Field data suggest that the proportion of mature individuals in the population is likely to be 0.4–0.6, indicating a total of 5,200–18,000 mature individuals, with no population known to have over 1,000 individuals (S. Hedges pers. comm. 2000). Most population figures given for numbers of Gaur are little more than guesses.
In Bangladesh, Gaur is probably extinct (Md Anwarul Islam in litt. 2008). A few were thought still to occur in the Chittagong, Chittagong Hill Tracts, Sylhet, and Mymensingh areas in the early 1980s (Sarker and Sarker 1984), but none had been seen in Pablakhali Wildlife Sanctuary in the Hill Tracts since the early 1970s (Khan 1985). Choudhury (2002) stated that Gaur individuals from Mizoram and Tripura (India) still cross into Bangladesh, although the records on which this was based were not given.
In Bhutan, Gaur apparently persists all over the southern foot-hill zone, notably in Royal Manas National Park, Phipsoo Wildlife Sanctuary and Khaling Wildlife Sanctuary (Choudhury 2002).
In Cambodia, Gaur probably declined by 90% or more in the period from the late 1960s to the early 1990s, at which time it still remained widespread, outside the lower Mekong and Tonle Sap lowlands, although in generally low numbers (Heng Kimchhay et al. 1998; Daltry and Momberg 2000; Timmins and Ou 2001; R. J. Timmins pers. comm. 2008). By far the most substantial Gaur population of the country remained in eastern Cambodia centred on Mondulkiri province, where in the late 1990s potentially several hundred to a thousand may have survived in a forested landscape of over 15,000 km² (Timmins and Ou 2001; Tordoff et al. 2005; R.J. Timmins pers. comm. 2008). Further declines, of about 50%, took place from the early 1990s to the present (R.J. Timmins pers. comm. 2008). As of 2007 the most significant population by far in Cambodia still remained within Mondulkiri Province (R. J. Timmins pers. comm. 2008 based on WCS and WWF unpublished data; T. Clements pers. comm. 2007; T.D. Evans pers. comm. 2007). Protection efforts in part of this landscape have probably stabilised the population there (R. J. Timmins pers. comm. 2008 based on WCS and WWF unpublished data; T. Clements pers. comm. 2007; T.D. Evans pers. comm. 2007). Protection outside these two areas is limited and for Cambodia as a whole the trend is still a significant decrease (R. J. Timmins pers. comm. 2008).
In China, Gaur occurs in Yunnan and southeast Tibet (Ma Yiqing and Wang Yingxiang 1995). All sites in Yunnan (except Yingjiang) formed, until the 1950s–1960s, one large contiguous population, but this is now heavily fragmented. By the 1980s, the species was extinct in Lancang county, and the remaining animals were split into two populations: Xishuangbanna–Simao and Cangyuan (Wang Yingxiang verbally 1998 to Duckworth and Hedges 1998). Some sources suggest that a population of 600–800 existed in Yunnan Province in the mid-1990s, with the majority occurring in Xishuangbanna National Nature Reserve (Ma Yiqing and Wang Yingxiang 1995); other sources suggest that only small numbers of Gaur remain in Yunnan (along the border with Myanmar) and that hunting had led to the local extinction of Gaur in Xishuangbanna (Xiang and Santiapillai 1993).
In India, three major (Western Ghats, Central India and North-East) and two minor (Bihar and West Bengal) “Gaur conservation areas” have been identified, reflecting the remaining distribution (Sankar et al. 2000; Choudhury 2002). The Western Ghats and their outflanking hills in south India constitute one of the most extensive extant strongholds of Gaur, with good numbers in Wynaad – Nagarahole – Mudumalai – Bandipur complex (Ranjitsinh 1997). Recent distributional assessment surveys in the Karnataka part of the Western Ghats showed the presence of Gaur in more than 60% of the 22,000 km² landscape (K.U. Karanth and N.S. Kumar unpublished data). Ranjitsinh (1997) estimated 12,000–22,000 in India, while Choudhury (2002) suggested that there were about 23,000–34,000 Gaur in India, Bhutan, and Bangladesh; but the true number is simply not known, as shown by the wide ranges. Major populations have been reported in Nagarahole National Park (probably over 2,000), Manas Wildlife Sanctuary (not known; may not be large), Bhadra Wildlife Sanctuary (over 800), Melghat Tiger Reserve (perhaps 500), Bandipur National Park (over 2,000), Kanha National Park (probably under 200), Radhanagari Wildlife Sanctuary (not known; may not be large), Tadoba–Andhari Tiger Reserve (over 1,000), Periyar Tiger Reserve and adjoining forest complex (500–1000), Annamalai range complex (Parambikulum, Valparai, and Grass Hills) (500–700), Silent Valley and adjoining forest complex (500–1000), Agastyamalai forest complex (about 500), Biligirirangangswamy Wildlife Sanctuary and Malemahadeswara Hill range (over 1000), Anshi–Dandeli Tiger Reserve and adjoining forest mosaic (perhaps about 400), Kudremukh National Park and Someshwara Wildlife Sanctuary (about 200–400), Brahmagiri–Pushpagiri–Talakaveri Sanctuary, Mukurti National Park, Pench Tiger Reserve, Jaldapara Wildlife Sanctuary (‘a large population’), Chapramari Wildlife Sanctuary (‘a large population’), and Gorumara Wildlife Sanctuary (‘a large population’). Many other areas are known or suspected to hold small populations, but in many sites such small numbers are unlikely to be viable (Choudhury 2002). The foregoing figures were assigned by N.S. Kumar (pers. comm. 2008) and are based on a variety of sources and methods, and from most areas are for guidance only. An increasing rigour is being applied to large mammal population estimation in India and some of the figures here will warrant modification and future changes to the quoted populations should not necessarily be seen as indicating that a real change in numbers has occurred. In recent years Gaur has reportedly been exterminated from three protected areas, Thattekad Wildlife Sanctuary (Kerala), Bhandhavgarh (Madhya Pradesh) and Kanger Valley National Park (Madhya Pradesh) (Pasha et al. 2004). Information on population status from the states of Andhra Pradesh, Orissa and North-East is limited (Choudhury 2002); there are demonstrably very few Gaur in Kaziranga (Karanth and Nicholls 2000). Recent population trends appear to have been stable in well protected areas. The following densities have been estimated: Bhadra Tiger Reserve, 1.48 +/- 0.63 (SE) per km² (Jathanna et al. 2003); Pench (Madhya Pradesh) dry deciduous forest, 0.7 animals per km² (Karanth and Nichols 2000); Nagarahole moist deciduous forest, 9.6 animals per km², making it the second-most abundant ungulate there (Karanth and Sunquist 1992); Bandipur dry deciduous forest, 7.0 animals per km² (Karanth and Nichols 2000); Tadoba-Andhari dry deciduous forest, 1.8 animals per km² (Karanth and Kumar 2005); Melghat dry deciduous forest, 1.0 animals per km² (Karanth and Kumar 2005); Pench (Maharashtra) dry deciduous forest, 0.8 animals per km² (Karanth and Kumar 2005). No densities could be estimated, despite the use of suitable survey methodology, at Kanha or Namdapha, implying relatively low populations, and none was found during such surveys at Kaziranga or Ranthambore (Karanth and Nichols 2000); only small numbers persist in Kaziranga (N.S. Kumar pers. comm. 2008). The Bhadra density is low, reflecting poaching (using snares, dogs and shotguns) and livestock grazing. However, the Gaur population there is now steadily increasing with successful conservation interventions (K.U. Karanth and N.S. Kumar unpublished data). A study at Nagarahole National Park compared the fauna of an area which was only moderately hunted with a heavily hunted site: this found respective densities of six and two Gaurs per km² (Madhusudan and Karanth 2002).
In Lao PDR, Byers et al. (1995) stated that the national population was approximately 1,000. This number is perhaps possible (field records exist from many areas), but animals would have been widely dispersed and at very low density in the extensive forest tracts across the country, with perhaps fewer than six sites holding more than 50 animals (Duckworth and Hedges 1998; Duckworth et al. 1999; Steinmetz 2004; R.J. Timmins pers. comm. 2008). A large population of Gaur was found in Nakai–Nam Theun NPA and the adjoining Nakai Plateau in the mid 1990s, when signs were readily found and even the animals themselves seen directly (Duckworth 1998; Duckworth and Hedges 1998; Evans et al. 2000). Resurveys of both areas, although using comparable methodology only on the plateau, indicate massive declines in the subsequent decade, with, for example, fairly intensive camera-trapping not recording the species at all (Dersu 2007; Johnson and Johnston 2007). The last decade has seen major reductions (in general, probably of well over 50%), in many large mammal populations across Lao PDR and it is now likely that many of the populations listed in Duckworth and Hedges (1998) have become locally extinct (Timmins and Robichaud 2005; R.J. Timmins pers. comm. 2008).
In Malaysia, in 1994 the population was stated to be less than 500 individuals and it was thought to be declining (J. Brennan pers. comm. to Read et al. 1994; a number of other stated numbers from around that time, all of the same general order of a few hundred, and none derived allowing a quantification of error margin, are presented in Hedges in prep.), with then confirmed records from various sites, e.g. the Sungai Singgor area of Temenggor Forest Reserve, Hulu Perak (Ratnam et al. 1995). In the subsequent few years, the population collapsed, despite optimism through relatively stronger control of guns in Malaysia than elsewhere on the southeast Asian mainland, and a perception (e.g. Nash 1993) of relatively strict control of wildlife trade in Malaysia. Densities in Taman Negara were crudely estimated from surveys during 1999–2001 as 0, 3 and 22 Gaurs per 100 km² in three study sites (Kawanishi and Sunquist 2004). Although these authors adjudged large mammal poaching to be negligible in Taman Negara, high levels were confirmed there by Wan Shahruddin (1998) and camera-trapping (suitable in methodology and intensity to find Gaur if any significant numbers were present) did not record Gaur at a fourth sector of Taman Negara in 1999 (Lynam et al. 2007). No Gaur was photographed at all in eight other potential Tiger areas surveyed by Lynam et al. (2007) during 1997–1999, including Temenggor (see Ratnam et al. 1995); because these were selected as the areas most likely to support Tigers and as Tiger-oriented camera-trapping elsewhere in South-east Asia usually yields photographs of Gaur, this study proves that only very low numbers of Gaur remain in Malaysia, no doubt way below the estimate in Read et al. (1994). Further supporting this, none was found in the Jerangau Forest Reserve, from which it was suspected to have been eradicated by hunting (Azlan 2006). Gaur was found in only two of 14 sites (seven virgin jungle reserves and seven adjacent logged areas), making it among the most localised mammal species which was detected in a study assessing the importance of virgin jungle reserves. The two occupied sites were a contiguous pair in each management class, and among the biggest sites surveyed (over 100 km²); the virgin jungle reserves themselves rarely exceed 3 km² (Laidlaw 2000). In the 1960s, there were estimated to be approximately 40 Gaur in Krau Wildlife Reserve, a figure deemed optimistic at the time; in the 1980s there were thought to be 22–31; in 1995 the estimate was only seven. When surveying the reserve in 2000 (see DWNP/DANCED 2002), R. Dunlop (pers. comm. 2008) covered the whole reserve (about 440 km²) with camera-traps in 119 one km-grid squares. Out of a total of 6,918 periods of 24-hour trapping, 61 species were recorded in 1,804 photographs of wildlife, yet only one photograph was of a Gaur. In almost 100 days of field searching for signs, Gaur tracks were recorded only on eight days, on each occasion as prints of isolated animals. The photograph, the sole sighting, and the eight track records were all in the south-east lowland area of the reserve, moving between an area of salt-licks and grazing just outside the reserve. This survey corroborated the 1995 estimate of seven animals, and the 2000–2001 figure could be even lower. Sumatran Rhinoceros Dicerorhinus sumatrensis and Asian Elephant Elephas maximus have both disappeared from Krau, and Gaur may be the next species to go. R. Dunlop (pers. comm. 2008) considers there no reason to expect this recent history not to typify that of Gaur populations at all sites in Malaysia. There are very recent records from: Tanjong Mentong in Taman Negara, where tracks were found during elephant surveys in late 2006 (Nurul Huda, WCS Malaysia pers. comm. 2008); Belum, in 2007, where animals were camera-trapped at salt-licks several times, including a herd of seven (Ahmad Zafir, WWF Malaysia pers. comm. 2008); and Temenggor, where individual animals (no herds) were camera-trapped nine times (never at salt licks, always on logging roads) in 2,364 trap-nights in 25 locations (Mark Darmaraj, WWF Malaysia pers. comm. 2008).
In Myanmar, the status of Gaur is poorly known (J.W. Duckworth and Than Zaw pers. comm. 2006). A March 1994 survey of Taminthi Wildlife Sanctuary, Myanmar's largest protected area, guessed that 100–200 Gaur remained, and that they were in danger of being extirpated from the area if current levels of poaching continued (Rabinowitz et al. 1995). A national Tiger survey camera-trapped Gaur in 11 of 15 camera-trapped sites, commonly in several (Lynam 2003); these sites were selected for their chances of holding Tigers, so, because the two species face similar threats, it may present the most positive picture of Gaur occurrence in the country.
In Nepal, the Gaur population was estimated to be 250–350 in the mid-1990s, with the majority in Chitwan National Park and Parsa Wildlife Reserve; population trends appeared to be relatively stable (J. Heinen pers. comm. to S. Hedges 1995). A few other areas hold small numbers (Choudhury 2002).
In Thailand, an estimated total of 920 Gaur remained in 1994. Only 6–7 populations were thought to contain 50 or more animals and there were no Gaur outside protected areas. The most important protected areas for Gaur were Khao Yai National Park, Tap Lan National Park and Pang Sida National Park, Huai Kha Khaeng Wildlife Sanctuary, Thung Yai Naresuan Wildlife Sanctuary, Kaeng Krachan National Park and Mae Nam Phachi Wildlife Sanctuary, the Khlong Nakha complex, and possibly Om Koi and Mae Tuen Wildlife Sanctuaries. Even in protected areas with much suitable habitat Gaur densities were very low, for example in Huai Kha Khaeng Wildlife Sanctuary the population density was less than 50% of the estimated carrying capacity of the area (Srikosamatara and Suteethorn 1995; S. Hedges pers. comm. 2000). Gaur populations are probably now increasing in several sites in Thailand, notably in Huai Kha Khaeng Wildlife Sanctuary and Thung Yai Naresuan Wildlife Sanctuary, with effective tackling of poaching. In Thung Yai Naresuan Wildlife Sanctuary, sign monitoring in one small area (20 km²) over six years since poaching absolutely ceased indicated an annual population growth rate there of 0.31, although the relative contributions of birth and immigration could not be distinguished. Breeding herds became much more common, compared with the situation 10–15 years previously at the height of poaching (Steinmetz et al. in prep.). In the Khao Paeng Ma reforestation site adjacent to Khao Yai National Park the population apparently expanded from two to 100 Gaur in less than 10 years, initially reflecting recolonization but apparently mostly from reproduction among animals at the site, all within a 10 km² area of secondary forest. The population stabilized at about 100 animals during 2004–2007 (based on counts from hilltops). As the forest grows and browse and grass diminish, reproduction is likely to decline (R. Steinmetz pers. comm. 2008). This herd has flourished only by dint of the provision of 24-hour security (A.J. Lynam pers. comm. 2008). Numbers also seem to be expanding rapidly in Kuiburi National Park in southern Thailand, again from an artificially lowered population and again within a secondary forest area adjacent to semi-evergreen forest. In early 2008, a (temporary) herd of almost 50 animals was spotted by elephant-watchers in a tree hide; only a few years previously, there were probably far fewer than this in the whole area (Steinmetz et al. 2007; R. Steinmetz pers. comm. 2008). By contrast, Gaur is largely gone from southern Thailand forest remnants, although it may be safe on the Malaysian border, reflecting tight control of illegal access and activities by the Border Patrol Police, and the danger of operations because of insurgency (A.J. Lynam pers. comm. 2008).
In Viet Nam, the current status of Gaur is poorly known. Several areas in Dak Lak Province (the Easo and Easup areas and Yok Don National Park) were known to contain Gaur in 1997 (Le Xuan Canh et al. 1997) but very little reliable information was traced for other parts of the country by Duckworth and Hedges (1998). Several herds of Gaur persist in Cat Tien National Park and in adjacent state forest enterprises (Polet and Ling 2004). There seems little room for doubt that the Gaur population is in serious decline, and many of the sites reported in Duckworth and Hedges (1998) may now have lost the species.
|Habitat and Ecology:||
The Gaur occurs from sea level up to at least 2,800 m asl (Wood 1937; Wharton 1968; Choudhury 2002). Despite the many reports that call it an animal of hill-country, low-lying areas seem to comprise optimal habitat (Choudhury 2002): in Conry’s (1989) study area, elevations ranged from 46 to 1,079 m asl but the three radio-tracked Gaur only used areas below 381 m. Elevations below 61 m were used most intensively and all three animals selected these low-lying areas; elevations above 61 m were selected against or used in proportion to availability (Conry 1989). Similarly, in the Tenasserim–Dawna mountains, Thailand, signs of Gaur were more abundant in the lowlands than in the hills, noting that this was the opposite of the patterns that would be predicted if hunting (itself concentrated heavily in the lowlands) was the chief determinant of population densities, although solitary animals were found mainly in the hills (Steinmetz et al. 2008). Schaller (1967) compared vegetation maps of India with the (mid-20th century) distribution of Gaur and concluded that the species was largely confined to evergreen, semi-evergreen, and moist deciduous forest but it also occurred in dry deciduous forest areas at the periphery of its range. He considered Gaur habitat to be "characterized by (1) large, relatively undisturbed forest tracts, (2) hilly terrain below an altitude of 5,000 to 6,000 ft (1,500–1,800 m asl), (3) availability of water, and (4) an abundance of forage in the form of coarse grasses (including bamboo), shrubs, and trees". However, He cautioned that the apparent preference of Gaur for hilly terrain may be partly due to the earlier conversion of most of the plains and other low lying areas to croplands and pastures. Similarly, Wharton (1968) concluded that Gaur appeared to avoid evergreen rainforest, "preferring foothill tracts of sub-humid or deciduous forest adjacent to savanna forest, glades or other open terrain affected by man and fire thus co-existing with and exploiting low human populations in hill zones with moderate to heavy rainfall". Throughout the range of Gaur, habitat destruction and human presence is usually higher in lowlands than in hills, meaning that differential human pressures may have caused the apparent preference of Gaur for hills. Gaur occurs in many forest types in Lao PDR, Viet Nam and Cambodia including truly evergreen and montane forest, but populations in closed canopy forests especially those at higher elevations might naturally be at much lower densities than those in lowland areas with a mosaic of open and closed forest types, and in closed canopy forests areas the species appears to frequent grassy openings (Duckworth et al 1999; Greiser Johns 2000; Duckworth and Hedges 1998; Timmins and Ou 2001; R.J. Timmins pers. comm. 2008). No significant difference was found in Gaur densities in moist compared with in dry deciduous forests in Nagarahole, India (Karanth and Sunquist 1992). Steinmetz et al. (in prep.) considered that Gaur herds in an expanding population were consistently concentrated in mixed deciduous forest rather than semi-evergreen forest, although the total area of study, that was safe from poaching, was only 20 km².
Gaur can persist in fragmented areas with some habitat disturbance and hunting, including in landscapes dominated by plantations (such as on parts of the Valparai plateau, south India), but in Southeast Asia it has been too heavily hunted in such areas to survive in them. Gaur can better tolerate rugged terrain and denser forest than other wild and domestic cattle, provided that there are adequate water sources (N.S. Kumar pers. comm. 2008.; Smith and Xie Yan in press). In Thailand it uses former cultivation amid forest, and populations in such areas can recover well if not hunted (Anak Pattanavipool pers. comm. 2006; R. Steinmetz pers. comm. 2006). A study of coffee estates around Bhadra Wildlife Sanctuary, India, recorded Gaur only in those coffee areas within 1 km of the sanctuary’s boundary (Bali et al. 2007).
Gaur both grazes and browses, reportedly eating mostly young green grasses but also leaves, fruit, twigs, and bark of various woody species, as well as coarse dry grasses, and bamboo. It seems able to maintain good condition on relatively low quality feed. At least 180–190 species of plants have been recorded in the diet (N.S. Kumar pers. comm. 2008). More information is given about Gaur’s diet in Dunbar Brander (1923), Hubback (1937), Schaller (1967), Wharton (1968), Krishnan (1972), Khan (1973), Conry (1981), Ebil Yusof (1982), Ramachandran et al. (1986), Prayurasiddhi and Smith (1993) and Sankar et al. (2000). Captive Gaur are reported to eat about 20 kg of green fodder per day (Ramachandran et al. 1986). During the cool season (November–February) the Gaur that Schaller (1967) studied in central India ate various herbs, large quantities of the leaves and seeds of bamboo, various other grasses, and the leaves of a number of tree species; but by the beginning of the hot season (March–June) coarse dry and semi-dry grasses made up the bulk of their diet, averaging 85% by volume (range 66–100%; four rumen samples), with browse only making up about 10%. Gaur also feeds on crops including corn, cassava, and young rubber trees, sometimes causing great damage (Wharton 1968; National Research Council 1983). Schaller (1967) reported that Gaur in Kanha National Park drank at least once per day during the hot season, usually in the evening. Sahai (1972) stated that Gaur drink at least twice in a 24-hour period but there appears to be no fixed time for drinking.
In undisturbed areas, Gaur is mainly diurnal, but in areas of high human disturbance, it is reported to become mainly nocturnal.
The basic social group appears to be a female–juvenile unit with larger groups tending to be more-or-less temporary assemblages (as in the other large species of Asian Bovini). Maternal herds containing several adult cows, juveniles, and calves frequently occur but their composition is somewhat fluid, particularly during the rut; these herds can often contain several subadult and adult males. The number of bulls in a herd changes with the time of year. Mature males associate with cows primarily during the rut: during Schaller’s study in Kanha National Park 65% of herds were accompanied by a mature male during April (the peak of the rut) but during the second half of the year the proportion dropped to 31% (Schaller 1967). Neither Schaller (1967) nor Sahai (1972) found any evidence to support Powell’s (1964) statement that young bulls usually live apart from the maternal herds; nor did Schaller’s observations support Dunbar Brander’s (1923) statement that the ‘master bull’ drives all other males from the herd. Males not associating with mixed herds either live in all-male groups or are solitary. The bull herds are generally unstable, although sometimes two or more animals may frequently associate. The typical herd size appears to be in the range of 5–12 animals and herds are rarely larger than 20 (Dunbar Brander 1923; Hubback 1937; Hislop 1961a; Schaller 1967; Prater 1971; Sahai 1972; Pal and Guin 1986). Large groups of 20, 40, or even 100 animals are probably temporary aggregations of smaller groups (Pal and Guin 1986).
The majority of the 125–150 Gaur studied by Schaller (1967) in Kanha National Park (India) ranged over an area of at least 78 km²; and the herds he studied travelled on average 2–3 miles (3.2–4.8 km) a day, although on occasion the distance could be much less. In West Malaysia Gaur home ranges have been estimated to be: approximately 13 km² (the Kuala Tahan herd in Taman Negara National Park) (Weigum 1972); and 29.9 km² for a yearling male, 52.1 km² for a yearling female, and 137.3 km² for an adult male in Pahang (Conry 1989). The large differences between the two studies probably reflect the different methods employed by the two workers (see Conry 1989: 60). Home ranges that can be as large as 137 km² hinder the setting aside of conservation areas for Gaur, particularly as Gaur often occurs in those areas where the pressures for economic development are highest.
In central India, most Gaur mate in December and January, and calves are born in August and September according to Dunbar Brander (1923); Stebbing (1911) and Sanderson (1912) generally agreed with Dunbar Brander but noted that calves may also be born in April, May, and June. In Kanha National Park (central India) Schaller (1967) recorded rutting bulls from December to June, with an apparent peak of sexual behaviour in March and April; however, the occurrence of a few new-born animals throughout the year indicated that some mating must occur in every month. In southern India the majority of matings reportedly take place between November and March (Morris 1937). In Cambodia the majority of Gaur births occur in December and January (Wharton 1957). In Myanmar, calves are born throughout the year (Evans 1912; Peacock 1933); and in West Malaysia young Gaur are seen in all months except October to December (Hubback 1937).
The gestation period of Gaur is reported to be nine months by Hubback (1937) and Crandall (1964) and 270 days, ‘a little shorter than for Banteng or domestic cattle and longer than for yak and Kouprey’, by National Research Council (1983: 49). Only one calf is born to a cow per pregnancy; there are no records of twins (Hubback 1937; Schaller 1967). Females probably first mate when about two years old and thus have their first calf at three years old (Schaller 1967). If the food supply is favourable, most adult females probably calve every year. In Kanha National Park, approximately 90% of adult cows bore young during a year but calf mortality was high, with about 50% of the calves dying before reaching one year old; the main cause of death was predation by Tigers Panthera tigris (Schaller 1967).
Rarely, bulls kill each other when fighting, and occasionally they are severely injured while fighting. Predation is by far the major cause of Gaur mortality, perhaps followed by disease (N.S. Kumar pers. comm. 2008). For example in Bandipur, Gaur is the most important prey in terms of the biomass taken by Tiger (42%) and Leopard Panthera pardus (15%) (Andheria et al. 2007). In Bandipur, Gaur remains were found in about 24% of Tiger faeces and 9% of Leopard faeces (Andheria et al. 2007). In adjoining Nagarahole, the proportion of Gaur in Tiger’s diet was about 45% in kills, 17% in scats (Karanth and Sunquist 1995). In Tadoba-Andhari, Gaur together with Sambar contributed to nearly 70% of the prey biomass consumed by Tiger, whereas in Pench-Maharashtra it was 80% (Karanth and Kumar 2005). In well protected forest reserves where different body-sized ungulates are abundantly available, Gaur is selectively predated by Tiger (N.S. Kumar pers. comm. 2008).
The set of threats and the gravity of each threat to Gaur differs greatly between south and central India, and Southeast Asia; China and most of northeast India are more similar to Southeast Asia in this respect (Farshid Ahrestani pers. comm. 2008), and are so treated in the following discussion.
In Indochina, Myanmar, and northeast India, hunting is the major problem, compounded by loss of suitable habitat: large tracts of suitable habitat hold few or no Guar. Ongoing habitat degradation and conversion is continually reducing the potential population, should hunting be brought under control (Duckworth et al. 1999; Duckworth and Hedges 1998; Timmins and Ou 2001; Choudhury 2002; Steinmetz 2004; Than Zaw pers. comm. 2006; R.J Timmins pers. comm. 2008; Farshid Ahrestani pers. comm. 2008). Hunting is often for meat and a variety of methods is used (Choudhury 2002). There is also a flourishing international trade in Gaur parts, both horns (decorative) and internal organs (for medicinal purposes). Given the small size of the remaining Southeast Asian subpopulations and the number of trophies found for sale in Cambodia, Lao PDR, Thailand, and Myanmar during essentially opportunistic surveys, trade is clearly a major threat (Srikosamatara et al. 1992; Baird 1993; Srikosamatara and Suteethorn 1994, 1995; Martin and Phipps 1996; Srifa La-Ong et al. 1997; Duckworth et al. 1999). The most important population in Cambodia is scattered through a forest landscape that encompasses four provinces (Mondulkiri, Kratie, Stung Treng and Ratanakiri) and five conservation areas (Lomphat Wildlife Sanctuary, Phnom Prich Wildlife Sanctuary, Siema Biodiversity Conservation Area, Mondulkiri Protection Forest (including the Srepok Wilderness area) and Phnom Nam Lyr Wildlife Sanctuary). Hunting is still rife in much of this area, and forest fragmentation is rapidly accelerating with human population immigration, infrastructural developments (especially roads), commercial agricultural expansion, economic land speculation and mineral extraction. However, although perhaps less than 20% of this area is well protected from the aforementioned threats and protected area management is only close to effective in two areas, the most significant constraint to the area’s Gaur is the long-term uncertainty of continuation of effective conservation management of the Srepok Wilderness Area and Siema Biodiversity Conservation Area. Although conservation efforts for Gaur and many other species are, in both these areas, extremely encouraging, both face an uncertain future with the possibility of degazettement of conservation status of parts of them, the possible loss of adequate external funding necessary to maintain high standards of management, the possible loss of political support necessary to uphold high protection standards and the uncertainties of maintaining a motivated well trained staff. Almost throughout Southeast Asia, protected areas are not yet stalling declines in mammals with commercial value, such as Gaur. Areas with stabilized populations (in addition to the two Cambodian areas, perhaps only a few areas in Thailand (see Population) and Cat Tien National Park in Viet Nam) are exceptional (R. Steimetz pers. comm. 2008; R.J. Timmins pers. comm. 2008).
Surprisingly, given its retention of healthy populations of other high-value very large mammals (Tiger and Asian Elephant Elephas maximus; Lynam et al. 2007; S. Hedges pers. comm. 2008 for elephants), Malaysia has among the most rapidly declining populations of Gaur. This reflects targeted hunting for large mammals of high commercial value, even within protected areas (Abdul Kadir 1998; Wan Shahruddin 1998; Lynam 1999; DWNP/Danced 2002; Lynam et al. 2007). Given the rapid decline within the 1990s and the lack of information traced since 2002, the exact current situation of Gaur in Malaysia is unclear. Kemasul Forest Reserve, where the species was found in Laidlaw’s (2000) survey of virgin jungle reserves, probably no longer exists as such. Even at the time of survey, most of the Forest Reserve was due to be converted into an Acacia mangium plantation, although still nominally a Forest Reserve. Kemasul was, long ago, part of a massive block of lowland forest which included the famous Jengka Triangle and Krau "Game" Reserve. Only Krau remains, albeit much reduced (R. Dunlop pers. comm. 2008). Even Krau, which was set up (in 1923) principally for the protection of Gaur, numerous in the area then, faces multiple threats (R. Dunlop pers. comm. 2008). In the surveys for DWNP/DANCED (2002), evidence of hunting, including cartridges, was found within the reserve, particularly in the easily accessible lowland areas; there were plenty of hunting stories by and about all of the different local communities, without exception, although their preferred target species differed. Conversion from natural forest to other land-uses of adjacent state-land combined with new ease of access has undoubtedly made Krau more of a focal point for such activity. There is a new (metalled) road and land conversion right up to the boundaries, facilitating access and easing the use of hunting methods such as spotlighting, as well as reducing habitat. The salt licks (for which Krau used to be famous) increasingly fall outside the forest (reflecting shrinking reserve boundaries and farmland encroachment into the reserve) and are now mostly in zones of high human activity of one sort or another. Temporary grasslands generated through traditional indigenous farming, which were often fairly difficult to get to and thus relatively free from hunting, were previously important Gaur habitat. A forester who used to work in Malaysia in the 1940s said that Gaur herds used to follow the Orang Asli settlements, moving in when they moved out, and Weigum (1972) even recommended "...to maintain herds in the park it is recommended that 45 acres of land be clear-cut preferably in 15 acre plots ½ mile apart and be recut every three years". Gaur meat is, or at least was being, sold at restaurants in placed like Cameron and Grik.
In south and central India, densities are likely to be widely below ecological carrying capacity, through hunting and competition with domestic livestock, although this conclusion is based on rather few empirical studies and there are some outstanding populations in well-secured protected areas. Substantial declines and local extinctions of Gaur were driven by hunting (for meat), competition with livestock, and habitat conversion, particularly during the first half of the twentieth century, resulting in a drastically reduced and fragmented distributional range, especially outside the protected area network (N.S. Kumar pers. comm. 2008). Hunting is much less prevalent than in Southeast Asia, reflecting the general enforcement of wildlife protection laws (Farshid Ahrestani pers. comm. 2008). Nonetheless, even in high-profile reserves, it is still suppressing densities significantly. For example, densities in Bhadra Tiger Reserve are very low, because of poaching, by comparison with what the habitat should support (Jathanna et al. 2003), and the measured density in a heavily hunted area of Nagarahole National Park was only a third of that in a well secured area (Madhusudan and Karanth 2002). Even though all hunting is illegal in this protected area, preventive measures are uneven across it, and Gaur is still poached, at least sporadically, wherever protection effectiveness is low (N.S. Kumar pers. comm. 2008). In recent years Gaur, has reportedly been exterminated from three Indian protected areas, Thattekad Wildlife Sanctuary (Kerala), Bhandhavgarh (Madhya Pradesh) and Kanger Valley National Park (Madhya Pradesh) (Pasha et al. 2004). Interaction with domestic stock is greatly under-appreciated as a threat to Gaur, but is probably the main factor which currently limits populations in south and central India: livestock are widely grazed even in otherwise well-managed protected areas. Mean densities of Gaur were fully 132 times higher in livestock-free areas than in adjacent livestock-grazed areas of Bandipur, and in shared grazing areas Gaur densities declined sharply with increasing livestock densities; in the grazed area studied, halving the livestock density allowed increases of Gaur by a factor of 57 (Madhusudan 2004). Threats to southern India’s forest ungulates by competition with domestic stock grazing within protected areas are exacerbated where dung is collected for export to adjacent coffee areas. Fuel wood removal may also be at levels sufficient to disrupt nutrient cycles of the habitat (Madhusudan 2005). Many Indian forest areas are severely encroached by exotics such as Lantana camara, Parthenium spp. and Chromolaena odorata, and these are suspected to effect major changes to forest structure (Hiremath and Sundaram 2005): but the effects of these on Gaur populations warrant further study. Killing of Gaur in retaliation to crop damage (through consumption and trampling) occurs, but is of limited significance to conservation (Choudhury 2002; Farshid Ahrestani pers. comm. 2008). Some animals also die from ingestion of agrochemicals, and human response when wandering into farmland and villages; these could be significant for already isolated and reduced populations (Choudhury 2002).
Throughout the range, diseases, particularly rinderpest and foot-and-mouth disease, transmitted by domestic cattle are a potentially serious threat. Foot-and-mouth is the most frequent, but rinderpest has had a particularly dramatic impact in India and several sub-populations of Gaur were nearly destroyed as a result of the disease in 1968 (Krishnan 1972; Pal and Guin 1986; Choudhury 2002), while in Bhadra, it nearly wiped out Gaur populations in 1988–1989, when nearly 800 individuals died (Girish D.V. pers. comm. 2008). In north and central Myanmar anthrax was a major factor in the decline of the species (Salter 1983). Pasha et al. (2004) considered that “in fact no wild animal in India [is] so profoundly influenced by transmitted infections from domestic livestock as Gaur”. There is little research or other evidence to determine impacts of introgression with domestic oxen on Gaur populations, although such factors might be expected to be most serious in India, given the wide overlap of domestic stock with Gaur. Farshid Ahrestani (pers. comm. 2008) saw a Gaur herd of 20 individuals far from human habitation with one domestic ox in the Palni Hills, so it seems likely that introgression occurs to at least some extent.
Srikosamtara and Suteethorn (1995) drew attention to the pressing threats facing Gaur, placing it in the same league of trade-driven threat as better-known species such as Tiger and Asian Elephant, and emphasising the international dimension of trade. Subsequently, the population collapse in Malaysia seems to have been driven by hunting teams from Lao PDR and Thailand (as proven by language at hunting camps in Temenggor), co-opting local Malaysian associates (A.J. Lynam pers. comm. 2008). It is reasonable to expect that when returns in Malaysia drop (as they probably have done already) these teams may turn their attention to other countries’ Gaur populations.
Gaur is listed in CITES Appendix I, and is legally protected in all range states. It occurs in many protected areas, although in north-east India, Myanmar, China, Lao PDR, Viet Nam and Malaysia many such areas retain only tiny populations or have already lost the species.
South and central India have a well established network of wildlife refuges and protection activities, to a large extent, are effective in many of them (Farshid Ahrestani pers. comm. 2008). However, conservation-oriented research and interventions on Gaur are limited and seem unlikely to expand much in the short-term, meaning that the species benefits largely through general protection activities (particularly those focused around Tiger and Asian Elephant), while species-specific negative factors or worrying long-term trends are likely to go unmanaged and even unnoticed (Farshid Ahrestani pers. comm. 2008). The chief of these seems to be competitive interaction with grazing stock: Gaur densities were the most depressed of all ungulates surveyed by Madhusudan (2004), perhaps reflecting higher niche overlap with domestic cattle than shown by the others (chiefly deer). There is long-term monitoring of some Gaur populations, for instance Mudumalai, Bandipur, Bhadra and Nagarahole (Farshid Ahrestani pers. comm. 2008). Rinderpest has reportedly been eradicated from India (Choudhury 2002). Choudhury (2002) listed proposed aspects of protected area expansion and consolidation to slow further fragmentation of Gaur populations in southern and central India.
In some protected areas (recent examples being Bhadra and Kudremukh in Karnataka), formerly forest-dwelling human communities have resettled closer to markets, clinics, school and other services (Karanth and Karanth 2007). This expands habitat available to Gaur (that which was formerly fields and houses) but more significant is the removal of a source of hunters and the cessation of grazing competition with domestic stock (see Karanth et al. 2006). Such conservation interventions have helped recovery of Gaur populations in this region. Increasing development aspirations mean that out migration is likely from further areas, e.g. Mudumalai (Farshid Ahrestani pers. comm. 2008).
As the majority of the Gaur population is in South Asia, conservation activities are important there. It is also important that populations are maintained at least in several areas in Southeast Asia, to maintain Gaur presence through its historical range, to preserve Gaur genetic diversity and through its importance to the ecosystems within which it lives. Given the regional pattern of threats and current successes to date, implementing effective conservation interventions is considerably more challenging, and therefore a higher priority in the Southeast Asian region.
Gaur occurs mainly in protected areas in Southeast Asia, although Cambodia and Myanmar may both have large populations outside them and some may persist is such areas in Lao PDR (Duckworth 1996; Hansel 2004). In most protected areas of Lao PDR, Viet Nam and Cambodia which still retain Gaur, protected status offers little, if any, real protection to the species, exceptions being Cat Tien National Park in Viet Nam, the Srepok Wilderness Area and the Siema Biodiversity Conservation Area in Cambodia; in the latter two Cambodian areas, there are now even signs of population recovery (R.J. Timmins pers. comm. 2008), as there are at Huai Kha Khaeng Wildlife Sanctuary and Thung Yai Naresuan Wildlife Sanctuary, Thailand, with effective tackling of poaching (R. Steinmetz pers. comm. 2008). Huai Kha Khaeng Wildlife Sanctuary and Thung Yai Naresuan Wildlife Sanctuary form part of Thailand’s ‘Western Forest Complex’ which if these recent gains are consolidated will, in the long term, hold the largest functional Gaur population in Thailand and perhaps South-east Asia.
The most important conservation measure for the species in Cambodia is the continued strengthening and support of conservation efforts in Siema Biodiversity Conservation Area and the Srepok Wilderness Area, and to a lesser extent support of protected area conservation management in the northern plains and central Cardamoms. Likewise in Viet Nam, the most important conservation measure is the continued support of protection efforts within Cat Tien National Park, in Thailand it is to continue the consolidation of the ‘Western Forest Complex’ and to continue the high-intensity protection of the rebuilt herd in Khao Paeng Ma reforestation site, adjacent to Khao Yai National Park. In Myanmar the Hukaung Valley is an exceptional area of lowland plains forest, grassland, and wetlands. Part lies within the Wildlife Sanctuary and some of the rest of the outstanding plains habitat is within an enormous proposed extension. Through its size and mostly little-encroached condition, this is the most outstanding remaining landscape-level floodplains habitat for very large mammals remaining in tropical Asia, and although wildlife populations are highly depleted, warrants the strongest effort to conserve it. It faces a number of daunting challenges (J.W. Duckworth pers. comm. 2008). In Lao PDR, any of a large number of areas could become key Gaur conservation sites, but there is as yet no precedent in the country for effective conservation of high-trade-value large mammals. Xe Pian, Nam Et–Phou Louey, and Nakai–Nam Theun NPAs could all be highly significant areas for the species (Duckworth and Hedges 1998; Steinmetz 2004; Johnson et al. 2006) but so could almost any other NPA; the reality is that the most success with Gaur conservation is likely to come through identifying areas with positive underlying situations to achieve conservation, rather than identifying the area with the most intrinsic importance for Gaur as the focus for efforts. Choudhury (2002) listed aspects of protected area expansion and consolidation to slow further fragmentation of Gaur populations in north-east India, Nepal, and Bhutan, and perhaps allow repopulation of Bangladesh.
The most important actions in Malaysia are to determine if any viable population remains in the country and, if so, to implement effective anti-poaching measures and to work at the policy level to allow effective control of large mammal hunting and trading. Under the current circumstances, any site-based activities are likely to require the sort of 24-hour security employed at the reforestation site adjacent to Khao Yai National Park, Thailand, which has allowed a rapid population growth of Gaur. It may also require reintroduction. The Department of Wildlife and National Parks have a breeding population of captive Gaur in enclosures in the Jenderak ranger station (within Krau Wildlife Reserve), for captive release.
A large captive population exists of wild-type Gaur (as well as of Mythun); given the complexity of field conservation of large ungulates, particularly in South-east Asia, it is clearly in resolving issues of hunting, trade, and protected area effectiveness that conservation interventions must focus.
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|Citation:||Duckworth, J.W., Steinmetz, R., Timmins, R.J., Pattanavibool, A., Than Zaw, Do Tuoc & Hedges, S. 2008. Bos gaurus. In: IUCN 2013. IUCN Red List of Threatened Species. Version 2013.2. <www.iucnredlist.org>. Downloaded on 21 April 2014.|
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