|Scientific Name:||Ursus thibetanus|
|Species Authority:||G. [Baron] Cuvier, 1823|
Seven subspecies have been recognized, many of which have been corroborated as distinct genetic clades (Kim et al. 2011, Yusefi 2013, Wu et al. 2015).
|Red List Category & Criteria:||Vulnerable A2cd ver 3.1|
|Assessor(s):||Garshelis, D. & Steinmetz, R.|
|Reviewer(s):||Obbard, M. & McLellan, B.|
|Contributor(s):||LaBruna, D., Abbas, F., Bendixsen, T., Broadis, N., Choudhury, A., Fahimi, H., Galbreath, G., Ghadirian, T., Hameed, S., Htun, S., Hwang, M., Islam, M., Jeong, D., Khan, M., Koike, S., Liu, F., Long, B., Ngoprasert, D., Oi, T., Olsson, A., Robinson, J., Sathyakumar, S., Scotson, L., Seryodkin, I., Shepherd, C., Wangchuk, S., Yadav, B., Yamazaki, K. & Yusefi, G.|
Widespread illegal killing of bears and trade in parts, combined with loss of habitat indicate that this species is likely declining in most parts of its range. Country bear experts on the IUCN SSC Bear Specialist Group (representing all range countries except the Democratic People's Republic of Korea) estimated rates of population change for the past 30 years (three bear generations), and projected rates of change for the next 30 years (see Figure 2 in the attached Supplementary Material). These assessments were based on perceived levels of exploitation, loss and degradation of habitat, and changes in area of occupancy within their respective countries; no range countries have estimates of abundance or indices of abundance that are sufficient to document population trend. One country (Pakistan) obtained empirical estimates of a decline in occupancy of 33% in 30-40 years (Abbas et al. 2015).
|Previously published Red List assessments:|
Fossil remains of the Asiatic Black Bear have been found in various sites in Europe, as far north as the Ural Mountains and Germany and west to France, dating from the early Pliocene to late Pleistocene (Erdbrink 1953, Kosintsev 2007, Baryshnikov and Zakharov 2013, Fourvel et al. 2014); however, in historic times the species has been limited to Asia. The western range limit is in southeastern Iran, inhabited by the so-called Baluchistan bear (U. t. gedrosianus) (Ahmadzadeh et al. 2008, Ghadirian et al. 2012). This small population is likely connected to the Baluchistan bear population in southern Pakistan. Disjunct populations of Asiatic Black Bears also occur in the more mountainous regions of northern Pakistan (Khan et al. 2012) and Afghanistan (Ostrowski et al. 2009). Eastward they continue within a narrow band along the foothills and south side of the Himalayas (up to treeline) across India, Nepal, and Bhutan, and then more widely distributed at lower elevations (generally >70 m but occasionally to 20 m) in the hill states of northeastern India (Sathyakumar and Choudhury 2007). They occur across mainland Southeast Asia, stretching south in Myanmar and Thailand to ~200 km north of the Malaysian border (Kanchanasakha et al. 2010); there are no records of Asiatic Black Bears ever existing in Malaysia. Over half the total range area of this species exists in China, especially in the south-central and southwestern parts of the country. This distribution includes portions of Tibet, from which the specific name, thibetanus, is derived. Smaller, remnant populations occur in eastern China. Another population cluster exists in northeastern China, the southern Russian Far East, and North Korea. A small isolated population exists in southern South Korea. They also live on the southern islands of Japan (Honshu and Shikoku) and on Taiwan and Hainan. Although they have been extirpated from large portions of their range, they remain in all 18 historic range countries.
Native:Afghanistan; Bangladesh; Bhutan; Cambodia; China; India; Iran, Islamic Republic of; Japan; Korea, Democratic People's Republic of; Korea, Republic of; Lao People's Democratic Republic; Myanmar; Nepal; Pakistan; Russian Federation; Taiwan, Province of China; Thailand; Viet Nam
|Range Map:||Click here to open the map viewer and explore range.|
The only rigorous population estimates for this species have been produced in Thailand, based on mark–recapture with camera traps, using distinctive chest patterns (Ngoprasert et al 2010; Higashide et al. 2012, 2013) to identify individual bears. This work yielded density estimates in three study sites in Khao Yai National Park, ranging from 8-29 bears per 100 km² (Ngoprasert et al. 2012, 2013). Bear sign density was found to be correlated with estimates of bear density, so sign surveys conducted across the country may, in the future, yield a reasonably reliable country-wide population index and trend information.
|Current Population Trend:||Decreasing|
|Habitat and Ecology:|
Asiatic Black Bears occupy a variety of forested habitats, both broad-leaved and coniferous, from near sea level to an elevation of 4,300 m (in northeastern India and Sikkim; Sathyakumar and Choudhury 2007, Sathyakumar et al. 2011). They also infrequently use open alpine meadows. A photo-capture was made in the alpine region of Nanda Devi Biosphere Reserve, Uttarakhand, India at 4,500 m (>1,000 m above the mean tree line; S. Sathyakumar, Wildlife Institute of India, pers. comm., 2014). In some areas of Nepal, local people have reported Asiatic Black Bears at higher than normal elevations, possibly a result of climate change (Aryal et al. 2012).
|Generation Length (years):||10|
|Use and Trade:||
Bear bile has been an important component of traditional medicine in Eastern Asia for millennia. The first written account of such use was recorded in the first pharmacopeia of China in 659 A.D. (Feng et al. 2009). Bear bile is used in Traditional Chinese Medicine (TCM) for reducing fever and inflammation, detoxifying the liver, arresting convulsions, improving eyesight, and dissolving gall stones.
Habitat loss due to logging, expansion of agriculture and plantations, roadway networks and dams, combined with hunting for skins, paws and especially gall bladders are the main threats to this species.
Protection of forested habitats would be an important conservation measure for this species. China, Thailand, and Viet Nam have imposed various sorts of logging bans, but with varying effects (Durst et al. 2001). In some cases this has resulted in trees being obtained (often illegally) from neighbouring countries, or in creating plantations which do not provide food resources for bears. However, in 2010 Russia banned the felling of Korean Pine, a key bear food source (I. Seryodkin, Russian Academy of Sciences, pers. comm. 2014).
|Errata reason:||The Supplementary Material was accidentally left off the assessment when it was published; this omission is corrected in this version of the assessment. Some country names have also been updated and a few typographical errors have been corrected.|
Abbas, F., Bhatti, Z.I., Haider, J., and Mian, A. 2015. Bears in Pakistan: distribution, population biology and human conflicts. Journal of Bioresource Management 2: 1-13.
Ahmadzadeh F., Liaghati, H., Kiabi, B.H., Mehrabian, A.R., Abdoli, A., and Mostafavi, H. 2008. The status and conservation of the Asiatic black bear in Nikshahr County, Baluchistan District of Iran. Journal of Natural History 42: 2379–2387.
Aramilev, V. V. 2006. The conservation status of Asiatic black bears in the Russian Far East. Understanding Asian Bears to Secure Their Future, pp. 86-89. Japan Bear Network, Ibaraki, Japan.
Aryal, A., Raubenheimer, D., Sathyakumar, S., Poudel, B. S., Ji, W., Kunwar, K. J., Kok, J., Kohshima, S., and Brunton, D. 2012. Conservation strategy for brown bear and its habitat in Nepal. Diversity 4(301-317).
Bargali, H.S. 2012. Distribution of different species of bears and status of human-bear conflict in the State of Uttarakhand, India. Advances in Biological Research 6(3): 121-127.
Baryshnikov, G.F., and Zakharov, D.S. 2013. Early Pliocene bear Ursus thibetanus (Mammalia, Carnivora) from Priozernoe locality in the Dniester Basin (Moldova Republic). Proceedings of the Zoological Institute of the Russian Academy of Sciences 317: 3-10.
Bista, R., and Aryal, A. 2013. Status of the Asiatic black bear Ursus thibetanus in the southeastern region of the Annapurna Conservation Area, Nepal. Zoology and Ecology.
Boatright, J.H., Moring, A.G., McElroy, C., Phillips, M.J., Do, V.T., Chang, B., Hawes, N.L., Boyd, A.P., Sidney, S.S., Stewart, R.E., Minear, S.C., Chaudhury, R., Ciavatta, V.T., Rodrigues, C.M.P., Steer, C.J., Nickerson, J.M., and Pardue, M.T. 2006. Tool from ancient pharmacopoeia prevents vision loss. Molecular Vision 12: 1706–1714.
Bromlei, F. G. 1965. Bears of the South Far-eastern USSR. Indian National Scientific Documentation Center, New Delhi, India.
Burgess, E.A., Stoner, S.S. and Foley, K.E. 2014. Brought to bear: an analysis of seizures across Asia (2000-2011). TRAFFIC Southeast Asia, Petaling Jaya, Selangor, Malaysia.
Carr, M. M., Yoshizaki, J., van Manen, F. T., Pelton, M. R., Huygens, O. C., Hayashi, H. and Maekawa, M. 2002. A multi-scale assessment of habitat use by asiatic black bears in central Japan. Ursus 13: 1-9.
Chang, H-C., Chang, H-J., Chao, T-Y. and Chan, S-Y. 1995. A survey of bear gallbladder commodities in the Taiwan market. Department of Health, Taipei, Taiwan.
Chape, S. Spalding, M. Jenkins, M. 2008. The World's protected areas: status, values and prospects in the 21st century. University of California Press, Berkeley, USA.
Charoo, S.A., Sharma, L.K., and Sathyakumar, S. 2011. Asiatic black bear-human interactions around Dachigam National Park, Kashmir, India. Ursus 22: 106–113.
Chongsomchai, Y. 2013. Habitat utilization and daily activity of Asiatic black bear (Ursus thibetanus) in Huai Kha Khaeng wildlife sanctuary. Kasetsart University, Bangkok.
Choudhury, A. 2011. Records of Black-breasted Parrotbill Paradoxornis flavirostris from Manas National Park, Assam, in north-east India. Forktail 27: 106-107.
Choudhury, A.U. 2013. Records of Asiatic black bear in North East India. Final report to International Association for Bear Research and Management (IBA). The Rhino Foundation for nature in NE India, Guwahati, Assam, India.
Choudhury, A.U. and Chand, S. 2012. Bear conservation action plan Assam. In: Sathyakumar, S., R. Kaul, N.V.K. Ashraf, A. Mookherjee, and V. Menon (eds), National bear conservation and welfare action plan, pp. 93-101. Ministry of Environment and Forests, Wildlife Institute of India, and Wildlife Trust of India.
Dar, N.I., .Minhas, R.A., Zaman, Q., and Linkie, M. 2009. Predicting the patterns, perceptions and causes of human–carnivore conflict in and around Machiara National Park, Pakistan. Biological Conservation 142: 2076–2082.
Durst, P.B, Waggener, T.R., Enters, T. and Cheng, T.L. 2001. Forests out of bounds: impacts and effectiveness of logging bans in natural forests in Asia-Pacific. Food and Agricultural Organization of the United Nations, Regional Office for Asia and the Pacific, Bangkok, Thailand.
Engin, F., Yermalovich, A., Ngyuen, T., Hummasti, S., Fu, W., Eizirik, D.L., Mathis, D., and Hotamisligil, G.S. 2013. Restoration of the unfolded protein response in pancreatic b cells protects mice against Type 1 diabetes. Science Translational Medicine 5.
ENV (Education for Nature Vietnam). 2012. New from the front line newsletter. Issue 2. Education for Nature Vietnam, Hanoi, Vietnam.
Erdbrink, D.P. 1953. A review of fossil and recent bears of the Old World with remarks on their phylogeny based upon their dentition. Drukkerij Jan de Lange, Deventer, Netherlands.
Fahimi, H., Yusefi, G.H., Madjdzadeh, S.M., Damangir, A.A., Sehhatisabet, M.E., and Khalatbari, L. 2011. Camera traps reveal use of caves by Asiatic black bears (Ursus thibetanus gedrosianus) (Mammalia: Ursidae) in southeastern Iran. Journal of Natural History 45: 2363–2373.
FAO. 2006. Global forest resources 2005. Progress towards sustainable forest management, Food and Agriculture Organization of the United Nations, Rome, Italy.
Feng, Y., Siu, K., Wang, N., Ng, K-M., Tsao, S-W., Nagamatsu, T., and Tong, Y. 2009. Bear bile: dilemma of traditional medicinal use and animal protection. Journal of Ethnobiology and Ethnomedicine 5.
Foley, K.E., Stengel, C.J. and Shepherd, C.R. 2011. Pills, Powders, Vials and Flakes: the bear bile trade in Asia. TRAFFIC Southeast Asia, Petaling Jaya, Selangor, Malaysia.
Fourvel, J-B, Fosse, P., Brugal, J-P., Cregut-Bonnoure, E., Slimak, L., and Tournepiche, J-F. 2014. Characterization of bear remains consumption by Pleistocene large carnivores (Felidae, Hyaenidae, Canidae). Quaternary International: 339–340: 232–244.
Galbreath, G. J., Hean, S. and Montgomery, S. M. 2000. A new color phase of Ursus thibetanus (Mammalia: Ursidae) from Southeast Asia. Natural History Bulletin of the Siam Society 49: 107-111.
Galbreath, G.J., Hunt, M., Clements, T. and Waits, L.P. 2008. An apparent hybrid wild bear from Cambodia. Ursus 19: 85-86.
Garshelis, D.L. 2002. Misconceptions, ironies, and uncertainties regarding trends in bear populations. Ursus 13: 321-334.
Garshelis, D.L. 2006. Assessing the status of the world’s bears—what can we tell from the trade in bear parts? In: Williamson, D. (ed.). Proceedings of the 4th international symposium on the trade in bear parts. pp. 6–20. TRAFFIC, East Asia–Japan, Tokyo.
Garshelis, D.L. and Scotson, L. 2012. World conservation congress votes to curtail bear farming. International Bear News 21: 12-16.
Garshelis, D.L., Joshi, A.R., Smith, J.L.D. and Rice, C.G. 1999. Sloth bear conservation action plan. In: C. Servheen, S. Herrero, and B. Peyton (eds), Bears: status survey and conservation action plan, pp. 225-240. IUCN/SSC Bear and Polar Bear Specialist Groups, Gland, Switzerland and Cambridge, U.K.
Ghadirian, T., Ghoddousi, A., Abolghasemi, H., Riazifar, A., Nekoei G., and Dehghani, M. 2012a. First camera-trap photos of Asiatic black bears in the Bashagard Region of southeastern Iran. International Bear News 21(1): 15-17.
Ghadirian, T., Ghoddousi, A., Qashqaei, A.T., Soufi, M. and Abolghasemi, H. 2012b. What can Asiatic Black Bears find to eat at the western edge of their geographic range (southern Iran)? . International Bear News 21(3): 27–28.
Gong, J. and Harris, R.B. 2006. The status of bears in China. Understanding Asian bears to secure their future, pp. 50-56. Japan Bear Network, Ibaraki, Japan.
Gurung, B., Smith, J.L.D., McDougal, C,. Karki, J.B., and Barlow, A. 2008. Factors associated with human-killing tigers in Chitwan National Park, Nepal. Biological Conservation 141: 3069–3078.
Gutleb, B. and Ziaie, H. 1999. On the distribution and status of the Brown Bear, Ursus arctos, and the Asiatic Black Bear, U. thibetanus, in Iran. Zoology in the Middle East 18: 5-8.
Hashimoto, Y. 2002. Seasonal food habits of the Asiatic black bear (Ursus thibetanus) in the Chichibu Mountains, Japan. Mammal Study 27: 65–72.
Hashimoto, Y., Kaji, M., Sawada, H. and Takatsuki, S. 2003. Five-year study on the autumn food habits of the Asiatic black bear in relation to nut production. Ecological Research 18: 485-492.
Higashide, D., Miura, S., and Miguchi, H. 2012. Are chest marks unique to Asiatic black bear individuals? Journal of Zoology 288: 199–206.
Higashide, D., Miura, S., and Miguchi, H. 2013. Evaluation of camera-trap designs for photographing chest marks of the free-ranging Asiatic black bear, Ursus thibetanus. Mammal Study 38: 35–39.
Honda, T., Yoshida, Y., and Nagaike, T. 2009. Predictive risk model and map of human-Asiatic black bear contact in Yamanashi Prefecture, central Japan. Mammal Study 34: 77–84.
Horino, S. and Miura, S. 2000. Population viability analysis of a Japanese black bear population. Population Ecology 42: 37–44.
Huygens, O. C. and Hayashi, H. 2001. Use of stone pine seeds and oak acorns by Asiatic black bears in central Japan. Ursus 12: 47-50.
Huygens, O. C., Miyashita, T., Dahle, B., Carr, M., Izumiyama, S., Sugawara, T. and Hayashi, H. 2003. Diet and feeding habits of Asiatic black bears in the Northern Japanese Alps. Ursus 14: 236-245.
Huygens, O., Goto, M., Izumiyama, S., Hayashi, H., and Yoshida, T. 2001. Denning ecology of two populations of Asiatic black bears in Nagano prefecture, Japan. Mammalia 65: 417-428.
Hwang, M.-H. 2003. Ecology of Asiatic black bears and people-bear interactions in Yushan National Park, Taiwan. Thesis, University of Minnesota.
Hwang, M.-H. and Garshelis, D. L. 2007. Activity patterns of Asiatic black bears (Ursus thibetanus) in the Central Mountains of Taiwan. Journal of Zoology (London) 271: 203-209.
Hwang, M. H., Garshelis, D. L. and Wang, Y. 2002. Diets of asiatic black bears in Taiwan, with methodological and geographical comparisons. Ursus 13: 111-125.
Hwang, M-H., Garshelis, D.L., Wu, Y-H., and Wang, Y. 2010. Home ranges of Asiatic black bears in the Central Mountains of Taiwan: Gauging whether a reserve is big enough. Ursus 21: 81–96.
Hwang, M-H., Pan, Y-J., R-A. Lin, R-A. 2012. Distribution model and conservation action plan for Formosan black bears. Forestry Bureau, Taipei, Taiwan.
Islam, M.A., Uddin, M., Aziz, M.A., Muzaffar, S.B., Chakma, S., Chowdhury, S.U., Chowdhury, G.W., Rashid, M.A., Mohsanin, S., Jahan, I., Saif, S., Hossain, M.B., Chakma, D., Kamruzzaman, M. and Akter, R. 2013. Status of bears in Bangladesh: going, going, gone? Ursus 24: 83-90.
IUCN. 2016. The IUCN Red List of Threatened Species. Version 2016-3. Available at: www.iucnredlist.org. (Accessed: 07 December 2016).
IUCN. 2017. The IUCN Red List of Threatened Species. Version 2017-1. Available at: www.iucnredlist.org.
Izumiyama, S. and Shiraishi, T. 2004. Seasonal changes in elevation and habitat use of the Asiatic black bear (Ursus thibetanus) in the Northern Japan Alps. Mammal Studies 29: 1-18.
Japan Bear Network. 2011. The report of accidents causing injury or death by bears. Japan Bear Network, Ibaraki, Japan.
Jeong, D-H., Lee, B-K., and Kang, J-K. 2011. The status of Asiatic black bears transplanted into Jirisan National Park, South Korea. . International Bear News 20(3): 14-16.
Kanchanasakha, B, Tanhikorn, S., Vinitpornsawan, V., Prayoon, A., and Paengphupha, G. 2010. Status of large mammals in Thailand. Department of National Parks, Wildlife, and Plant Conservation, Bangkok, Thailand.
Kang, S. and Phipps, M. J. 2003. A question of attitude: South Korea's Traditional Medicine practitioners and wildlife conservation. TRAFFIC East Asia, Hong Kong.
Khan, M., Rosen, T., and Zahler, P. 2012. Status and conservation of Asiatic black bears in Diamer District, Pakistan. International Bear News 21(1): 13-14.
Kim,Y-K., Hong, Y-J., Min, M-S., Kim, K. S., Kim, Y-J., Voloshina, I., Myslenkov, A., Smith, G.J.D., Cuong, N.D., Tho, H.H., Han, S-H., Yang, D-H., Kim, C-B., and Lee, H. 2011. Genetic status of Asiatic black bear (Ursus thibetanus) reintroduced into South Korea based on mitochondrial DNA and microsatellite loci analysis. Journal of Heredity 102: 165-174.
Kishimoto, R. 2009. Status of the 2006 drastic rise of Asiatic black bear (Ursus thibetanus) intrusions into residential areas in Nagano Prefecture. In: Oi et al. (ed.), Biology of bear intrusions, pp. 35–39.
Koike, S. 2009. Fruiting phenology and its effect on fruit feeding behavior of Asiatic black bears. Mammal Study 34: 47–52.
Koike, S. 2010. Long-term trends in food habits of Asiatic black bears in the Misaka Mountains on the Pacific coast of central Japan. Mammalian Biology 75(17–28).
Koike, S., and Hazumi, T. 2008. Notes on Asiatic black bears denning in the Misaka Mountains, central Japan. Ursus 19: 80–84.
Koike, S., Kozakai, C., Nemoto, Y., Masaki, T., Yamazaki, K., Abe, S., Nakajima, A., Umemura, Y., and Kaji, K. 2012. Effect of hard mast production on foraging and sex-specific behavior of the Asiatic black bear (Ursus thibetanus). Mammal Study 37: 21–28.
Koike, S., Masaki, T., Nemoto, Y., Kozakai, C., Yamazaki , K., Kasai, S., Nakajima, A., and Kaji, K. 2010. Estimate of the seed shadow created by the Asiatic black bear Ursus thibetanus and its characteristics as a seed disperser in Japanese cool-temperate forest. Oikos 120: 280–290.
Koike, S., Morimoto, H., Goto, Y., Kozakai, C., and Yamazaki, K. 2008. Frugivory of carnivores and seed dispersal of fleshy fruits in cool-temperate deciduous forests. Journal of Forest Research 13: 215–222.
Koike, S., Nakashita, R., Naganawa, K., Koyama, M., and Tamura, A., 2013. Changes in diet of a small, isolated bear population over time. Journal of Mammalogy 94: 361–368.
Kosintsev, P. 2007. Late Pleistocene large mammal faunas from the Urals. . Quaternary International 160: 112–120.
Kozakai, C., Yamazaki, K., Nemoto, Y., Nakajima, A., Koike, S., Abe, S., Masaki, T., and Kaji, K. 2011. Effect of mast production on home range use of Japanese black bears. Journal of Wildlife Management 75: 867–875.
Kozakai, C., Yamazaki, K., Nemoto, Y., Nakajima, A., Umemura, Y., Koike, S., Goto, Y., Kasai, S., Abe, S., Masaki, T., and Kaji, K. 2013. Fluctuation of daily activity time budgets of Japanese black bears: relationship to sex, reproductive status, and hardmast availability. Journal of Mammalogy 94: 351–360.
Liu, F., McShea, W., Garshelis, D., Zhu, X., Wang, D., and Shao, L. 2010. Human-wildlife conflicts influence attitudes but not necessarily behaviors: Factors driving the poaching of bears in China. Biological Conservation 144(538–547).
Liu, F., McShea, W., Garshelis, D., Zhu, X., Wang, D., Gong, J., and Chen, Y. 2009. Spatial distribution as a measure of conservation needs: an example with Asiatic black bears in south‐western China. Diversity and Distributions 15: 649–659.
Livingstone, E., and Shepherd, C.R. 2016. Bear farms in Lao PDR expand illegally and fail to conserve wild bears. Oryx 50: 176–184.
Ministry of the Environment. 2011. A Heisei 22 fiscal year report for national survey on the natural environment – investigation of living status of the specific wild animals and investigation for establishing monitoring footing. Biodiversity Center, Minsitry of the Environment, Japan.
Mizukami, R. N., Goto, M., Izumiyama, S., Hayashi, H. and Yoh, M. 2005. Estimation of feeding history by measuring carbon and nitrogen stable isotope ratios in hair of Asiatic black bears. Ursus 16: 93-101.
Mukesh, L.K. Sharma, S.A. Charoo, and Sathyakumar, S. 2015. Conflict bear translocation: investigating population genetics and fate of bear translocation in Dachigam National Park, Jammu and Kashmir, India. PLoS ONE 10(8).
Narita, R., Sugimoto, A., and Takayanagi, A. 2006. Animal componets in the diet of Japanese black bears Ursus thibetanus japonicas in the Kyoto area, Japan. Wildlife Biology 12(375–384).
Ngoprasert, D., Reed, D.H., Steinmetz, R. and Gale, G.A. 2012. Density estimation of Asian bears using photographic capture–recapture sampling based on chest marks. Ursus 23: 117-133.
Ngoprasert, D., Steinmetz, R., and Gale, G.A. 2010. The use of chest marks to distinguish Asiatic black bear and sun bear individuals in Thailand. International Bear News 19(2): 18-20.
Ngoprasert, D., Steinmetz, R., and Gale, G.A. 2013. Estimating the abundance of Asiatic black bears and sun bears in Thailand. International Bear News 22(3): 30-32.
Ngoprasert, D., Steinmetz, R., Reed, D.H., Savini, T., and Gale, G.A. 2011. Influence of fruit on habitat selection of Asian bears in a tropical forest. Journal of Wildlife Management 75: 588–595.
Nguyen Xuan Dang. 2006. The current status and conservation of bears in Vietnam. Understanding Asian bears to secure their future., pp. 61-65. Japan Bear Network, Ibaraki, Japan.
Oi, T., and Furusawa, H. 2008. Nutritional condition and dietary profile of Japanese black bear (Ursus thibetanus japonicas) killed in western Japan in autumn 2004. Mammal Study 33: 163–171.
Oi, T. and Yamazaki, K. 2006. The status of Asiatic black bears in Japan. Understanding Asian bears to secure their future, pp. 122-133. Japan Bear Network, Ibaraki, Japan.
Ostrowski, S., Zahler, P., Dehgan, A., Stevens, K., Karlstetter, M., and Smallwood, P. 2009. The Asiatic black bear still survives in Nuristan, Afghanistan. International Bear News 18(4): 14-15.
Phelps, J., Carrasco, L.R., and Webb, E.L. 2014. A framework for assessing supply‐side wildlife conservation. Conservation Biology 28: 244–257.
Rao, M., Myint, T., Zaw, T. and Htun, S. 2005. Hunting patterns in tropical forests adjoining the Hkakaborazi National Park, north Myanmar. Oryx 39(3): 292-300.
Reid, D., Jiang, M., Teng, Q., Qin, Z. and Hu, J. 1991. Ecology of the Asiatic black bear (Ursus thibetanus) in Sichuan, China. Mammalia 55: 221-237.
Rinzin, C., Vermeulen, W.J., Wassen, M.J., and Glasbergen, P. 2009. Nature conservation and human well-being in Bhutan: an assessment of local community perceptions. The Journal of Environment & Development 177–202.: 177–202.
Sakurai, R., and Jacobson, S.K. 2011. Public perceptions of bears and management interventions in Japan. Human–Wildlife Interactions 5: 123–134.
Sangay, T., and Vernes, K. 2008. Human–wildlife conflict in the Kingdom of Bhutan: patterns of livestock predation by large mammalian carnivores. Biological Conservation 141: 1272–1282.
Sathyakumar, S. 2006. The status of Asiatic black bears in India. Understanding Asian bears to secure their future, pp. 12-19. Japan Bear Network, Ibaraki, Japan.
Sathyakumar, S. 2006. The status of Asiatic black bears in India. Understanding Asian bears to secure their future . pp. 12-19. Japan Bear Network, Ibaraki, Japan.
Sathyakumar, S., and Choudhury, A. 2007. Distribution and status of the Asiatic black bear Ursus thibetanus in India. Journal of the Bombay Natural History Society 104: 316–323.
Sathyakumar, S., and Viswanath, S. 2003. Observations on food habits of Asiatic Black Bear (Ursus thibetanus) in Kedarnath Wildlife Sanctuary, India – Preliminary evidence on the role of black bears in enhancing seed germination and dispersal. Ursus 14(95–104).
Sathyakumar, S., Bashir, T., Bhattacharya, T. and Poudyal, K. 2011. Assessing mammal distribution and abundance in intricate eastern Himalayan habitats of Khangchendzonga, Sikkim India. Mammalia 75: 257-268.
Sathyakumar, S., Kaul, R., Ashraf, N.V.K., Mookherjee, A., and Menon, V. 2012. National Bear Conservation and Welfare Action Plan. Ministry of Environment and Forests, Wildlife Institute of India, and Wildlife Trust of India, India.
Saw Htun. 2006. The status and conservation of bears in Myanmar. Understanding Asian bears to secure their future, pp. 45-49. Japan Bear Network, Ibaraki, Japan.
Schaller, G. B., Teng, Q., Johnson, K. G., Wang, X., Shen, H. and Hu, J. 1989. The feeding ecology of giant pandas and Asiatic black bears in the Tangjiahe Reserve, China. In: J. L. Gittleman (ed.), Carnivore behavior, ecology, and evolution, pp. 212-240. Cornell University Press, Ithaca, NY, USA.
Scotson, L. 2010. Distribution and status of Asiatic black bear and Malayan sun bear in Nam Et Phou Louey National Protected Area, northeast Lao PDR. International Bear News 19: 14-16.
Scotson, L. 2012. Status of Asiatic black bears and sun bears in Xe Pian National Protected Area, Lao PDR. International Bear News 21: 8-11.
Scotson, L. and Brocklehurst, M. 2013. Bear poaching in Lao PDR is exposed as an increasing threat to wild populations. International Bear News 22: 22-23.
Scotson, L. and Hunt, M. 2012. Dismantling the “wall of death”: emergency bear snare-line patrol in the Nam Kan National Protected Area, Lao PDR. International Bear News 21: 17-19.
Scotson, L., Vannachomchan,K., and T. Sharp, T. 2014. More valuable dead than deterred: crop raiding bears in Lao PDR. Wildlife Society Bulletin 38: 783–790.
Servheen, C. 2013. Bear paw smuggling into China continues. International Bear News 22(3): 24.
Servheen, C., Herrero, S. and Peyton, B. 1999. Bears. Status Survey and Conservation Action Plan. IUCN/SSC Bear and Polar Bear Specialist Groups, Gland, Switzerland.
Seryodkin, I. V., Goodrich, J. M., Kostyria, A. V., Schleyer, B. O., Smirnov, E. N., Kerley, L. L. and Miquelle, D. G. 2005. Relationship between tigers, brown bears, and Himalayan black bears. In: D. G. Miquelle, E. N. Smirnov and J. M. Goodrich (eds), Tigers of Sikhote-Alin Zapovednik: ecology and conservation, pp. 156-163. PSP, Vladivostok, Russia.
Seryodkin, I. V., Kostyria, A. V., Goodrich, J. M., Miquelle, D. G., Smirnov, E. N., Kerley, L. L., Quigley, H. B. and Hornocker, M. G. 2003. Denning ecology of brown bears and Asiatic black bears in the Russian Far East. Ursus 14: 153-161.
Sharma, L.K., Charoo, S.A., and S. Sathyakumar, S. 2010. Investigations on the Ecology and Behaviour of Asiatic Black bear using Satellite Telemetry: A case study from the Dachigam National Park, Kashmir, India. Telemetry in Wildlife Science, ENVIS Bulletin: Wildlife & Protected Areas , pp. 86–94. Wildlife Institute of India, Dehradun.
Shepherd, C.R. and Nijman, V. 2008. The trade in bear parts from Myanmar: an illustration of the ineffectiveness of enforcement of international wildlife trade regulations. Biodiversity and Conservation 17: 35-42.
Solá, S., Garshelis, D.L., Amaral, J.D., Noyce, K.V., Coy, P.L., Steer, C.J., Iaizzo, P.A., and Rodrigues, C.M.P. 2006. Plasma levels of ursodeoxycholic acid in black bears, Ursus americanus: seasonal changes. Comparative Biochemistry and Physiology, Part C 143: 204–208.
Steinmetz, R. 2011. Ecology and distribution of sympatric Asiatic black bears and sun bears in the seasonally dry forests of Southeast Asia. Smithsonian Institution Scholarly Press, Washington, DC.
Steinmetz, R., Chutipong, W. and Seauturien, N. 2006. Collaborating to conserve large mammals in Southeast Asia. Conservation Biology 20: 1391–1401.
Steinmetz, R., Garshelis, D.L., Chutipong, W. and Seuaturien, N. 2011. The shared preference niche of sympatric Asiatic black bears and sun bears in a tropical forest mosaic. PloS ONE 6.
Steinmetz, R., Garshelis, D.L., Chutipong, W., and Seuaturien, N. 2013. Foraging ecology and coexistence of Asiatic black bears and sun bears in a seasonal tropical forest in Southeast Asia. Journal of Mammalogy 94: 1-18.
Steinmetz, R., Srirattanpon, S., Mor-Tip, J. and Seuaturien, N. 2014. Can community outreach alleviate poaching pressure and recover wildlife in Southeast Asian protected areas? . Journal of Applied Ecology 51: 1469-1478.
Stubblefield, C.H., and Shrestha, M. 2007. Status of Asiatic black bears in protected areas of Nepal and the effects of political turmoil. Ursus 18: 101–108.
Takahata C, Nielsen, S.E., Takii, A. and Izumiyama, S. 2014. Habitat selection of a large carnivore along human-wildlife boundaries in a highly modified landscape. PLoS ONE 9(1).
TRAFFIC. 2014. China’s business community leads the way in stopping the gifting of threatened species products.
Vang, S., Longley, K., Steer, C.J., and Low, W.C. 2014. The unexpected uses of Urso- and Tauroursodeoxycholic Acid in the treatment of non-liver diseases. Global Advances in Health and Medicine 3: 58–69.
Vu, Q.T. 2010. An analysis of attitudes and bear bile use in Vietnam. Education for Nature, Vietnam (ENV), Hanoi.
Wang, S.W., Lassoie, J.P., and P.D. Curtis, P.D. 2006. Farmer attitudes towards conservation in Jigme Singye Wangchuck National Park, Bhutan. Environmental Conservation 33(148–156).
Wen, C., and Wang, D. 2013. Update the status of Sun Bear in Yunnan, China. Unpublished report. International Association for Bear Research and Management.
Wu, J., Kohno, N., Mano, S., Fukumoto, Y., Tanabe, H., Hasegawa, M., and Yonezawa, T. 2015. Phylogeographic and demographic analysis of the Asian black bear (Ursus thibetanus) based on mitochondrial DNA. PLoS ONE 10(9).
Yadav, B.P., Aryal, A., Poudel, D.P., Pandey, P., and Thapalia, B.P. 2012. Asiatic black bear killed in Rara National Park, Nepal. International Bear News 21(3): 20-22.
Yamada, A., and Fujioka, M. 2010. Features of planted cypress trees vulnerable to damage by Japanese black bears. Ursus 21: 72–80.
Yamamoto, T., Oka, T., Ohnishi, N., Tanaka, H., Takatsuto, N., and Y. Okumura. 2012. Genetic characterization of northernmost isolated population of Asian black bear (Ursus thibetanus) in Japan. Mammal Study 37(85–91).
Yamanaka, A., Kiyoshi, Y., Tsunenori, T., Toshiki, M., Oi, T., Seigo, K., Shimozuru, M., and Tsubota, T. 2011. Estimating the success rate of ovulation and early litter loss rate in the Japanese black bear (Ursus thibetanus japonicus) by examining the ovaries and uteri. Japanese Journal of Veterinary Research 59: 31–39.
Yamazaki, K. 2003. Effects of pruning and brush clearing on debarking within damaged conifer stands by Japanese black bears. Ursus 14: 94-98.
Yamazaki, K, Kozakai, C., Koike, S., Morimoto, H., Goto, Y., and Furubayashi, K. 2012. Myrmecophagy of Japanese black bears in the grasslands of the Ashio area, Nikko National Park, Japan. Ursus 23: 52–64.
Yusefi G.H. 2013. Noninvasive genetic tracking of Asiatic black bears (Ursus thibetanus) at its range edge in Iran — a pilot study. M.S. Thesis. Uppsala University, Sweden.
|Citation:||Garshelis, D. & Steinmetz, R. 2016. Ursus thibetanus. (errata version published in 2017) The IUCN Red List of Threatened Species 2016: e.T22824A114252336.Downloaded on 26 July 2017.|
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