|Scientific Name:||Neophocaena asiaeorientalis (Pilleri & Gihr, 1972)|
|Infra-specific Taxa Assessed:|
The recognition of two externally distinct morphological forms of Finless Porpoises as separate biological species (Neophocaena phocaenoides and N. asiaeorientalis) was accepted when it was demonstrated that the two forms are reproductively isolated (and likely have been separated since the last glacial maximum) even though they are sympatric in a fairly large area of East Asia (Wang et al. 2008, Jefferson and Wang 2011). Differences in the external morphology of the dorsal aspect of the two species are distinguishable even amongst free-ranging animals (as opposed to animals in-hand for close inspection) (Wang et al. 2010) and intermediate individuals have never been reported even though several hundred to thousands of carcasses have been examined. The two species also clearly differ in craniometry (Amano et al. 1992, Jefferson 2002). Within the Narrow-ridged species, two subspecies are retained: the Yangtze Finless Porpoise, N. a. asiaeorientalis, and the East Asian Finless Porpoise or Sunameri, N. a. sunameri (Gao 1991, Wang 1992a,b; Gao and Zhou 1995). Where Narrow-ridged Finless Porpoises have been studied fairly well (e.g., Yoshida et al. 1995, 2001; Yoshida 2002, Jefferson 2002, Zheng et al. 2005, Yang et al. 2008, Chen et al. 2010, Xu et al. 2010, Li et al. 2011, Ju et al. 2012, L. Li et al. 2013, S. Li et al. 2013, Chen et al. 2014, Jia et al. 2014, Lin et al. 2014), there is evidence to suggest subpopulation structure.
|Red List Category & Criteria:||Endangered A2bcde+3bcde+4bcde ver 3.1|
|Assessor(s):||Wang, J.Y. , Reeves, R.|
|Reviewer(s):||Taylor, B.L., Sutaria, D., Minton , G., Amano, M., Jefferson, T.A. & Ponnampalam, L.|
Although the data are insufficient to make a rigorous quantitative assessment of population trend for N. asiaeorientalis throughout its range, the two subpopulations with suitably long time-series of abundance estimates both declined by >50% in far less than three generations and therefore easily meet criterion A for Endangered (see below). Another recent decline of approximately 70% in a large part of the species’ range (Yellow Sea, Korea) provides an index of decline (subcriterion b) that may apply throughout many unsurveyed portions of the range. Although the rate of decline in such areas can only be inferred, there is no evidence that threats differ substantially. The scale of threats is large enough over enough of the range to suspect and infer a decline of at least 50% over the last three generations (45 years; Moore 2015) (A2), over three generations into the future (A3), and over a three-generation period that includes both the past and the future (A4). The factor most responsible for such decline is incidental mortality in fisheries (subcriterion d), but the loss and degradation of habitat (including chemical pollution; subcriteria c and e) and vessel strikes (at least in the Yangtze River system; Turvey et al. 2013) are contributing factors as well. The primary causes of the suspected/inferred decline in population size—bycatch in fishing gear and mortality from vessel strikes (both interpreted here as “exploitation”), decline in habitat quality, and possibly pollution—have not ceased, may not be well understood, and may not all be reversible.
As the Scientific Committee of the International Whaling Commission concluded after a review of the species (at the time with no recognized species-level distinction between N. phocaenoides and N. asiaeorientalis) in 2005 (IWC 2006), “human populations adjacent to the Finless Porpoise’s habitat are increasing in size and becoming more industrialised so the expectation should be that anthropogenic pressures will continue and intensify.” None of the threats has been seriously addressed or mitigated in any part of the Narrow-ridged Finless Porpoise’s range, even though threat levels are likely increasing. Partly because of their small size, phocoenids are exceptionally vulnerable to incidental mortality in gillnets (e.g., Jefferson and Curry 1994). Incidental mortality in fishing gear is either known or presumed to occur throughout the range of Narrow-ridged Finless Porpoises (Reeves et al. 1997, IWC 2006).
There is clear evidence of a declining trend in three major parts of this species’ range. In the Seto Inland Sea of Japan, a decline of nearly 70% was estimated over a period of 22 years, from 1976-1978 to 1999-2000 (Kasuya et al. 2002, also see Shirakihara et al. 2007). There is also evidence of a rapid decline in recent decades in the Yangtze River and adjoining lake systems of China (Zhao et al. 2008, Wang 2009, Zhao et al. 2012) with a 60% decline in the six years between 2006 and 2012 (Mei et al. 2014); the subspecies there (N. a. asiaeorientalis) was classified as Endangered in 1996 and uplisted to Critically Endangered in 2013 (Wang et al. 2013). Finally, there is recent evidence suggesting a decline of approximately 70% in abundance of Narrow-ridged Finless Porpoises in the Korean portion of the Yellow Sea between the early 2000s and 2011 (Park et al. 2015), and there is no reason to believe this decline has been reversed.
There is sufficient information for separate assessment of at least two threatened subpopulations – the subpopulation in the Seto Inland Sea of Japan, which likely qualifies for Endangered or Critically Endangered (Hashimoto et al. 2015), and the Ariake Sound/Tachibana Bay (Japan) subpopulation, which likely qualifies for at least Vulnerable (Hashimoto et al. 2015).
|Previously published Red List assessments:|
The type specimen was from the Yangtze River, Kiangsu [=Jiangsu] Province, China, ~129 km (80 miles) northwest of Shanghai (Pilleri and Gihr 1972).
The distirbution map shows where the species may occur based on oceanography and known habitat preferences. Given the paucity of dedicated cetacean survey effort in much of the species’ suspected range, the distribution map includes not only the known range of the species but also the possible or projected range where oceanic features are consistent with the its known preferred habitat. As such, the species has not been recorded for all the states within the hypothetical range as shown. States for which confirmed records of the species exist are indicated in the occurrence list "Extant." States within the hypothetical range, but for which no confirmed records exist, are indicated as "Presence Uncertain."
Native:China; Japan; Korea, Republic of; Taiwan, Province of China
|FAO Marine Fishing Areas:|
Pacific – northwest
|Range Map:||Click here to open the map viewer and explore range.|
Narrow-ridged Finless Porpoises are small and cryptic and therefore difficult to survey. Estimates of abundance have been made for only a few areas (IWC 2006). Estimates from the early 2000s are available for five subpopulations in Japanese waters, distinguished on the basis of skull morphology and mtDNA variability (Yoshida et al. 1995, Yoshida 2002), as follows: 3,807 (coefficient of variation (CV) 16%) in Ariake Sound/Tachibana Bay (Shirakihara and Shirakihara 2002); 289 (CV 19%) in Omura Bay (Shirakihara and Shirakihara 2002); 3,743 (CV 24%) in Ise/Mikawa Bay (Yoshida 2002); 3,387 (CV 33%) in Chiba/Sendai Bay (Amano et al. 2003); and 7,572 (CV 17%) in the Inland Sea (Shirakihara et al. 2007). Most of the estimates were summarized in IWC (2006). For the Inland Sea subpopulation, Kasuya et al. (2002) recorded a decline in sighting rate of 18-100% for 18 tracklines surveyed in 1976–78 and again in 1999–2000. The declines for 11 of the tracklines were statistically significant. Using abundance indices calculated by multiplying the density indices (no. individuals/cruise distance) and area size of each stratum, an overall decline of 69% is inferred over the 22 years between the two sets of surveys (T. Kasuya pers. comm. 2007, data from Kasuya et al. 2002).
|Current Population Trend:||Decreasing|
|Habitat and Ecology:|
Narrow-ridged Finless Porpoises are found mainly in coastal waters, including shallow bays, possibly mangrove swamps, estuaries, and some large rivers. However, they can also occur in shallow waters (<200 m deep) quite far from shore (up to 240 km). They appear to have a strong preference for waters with a sandy or soft bottom (Jefferson and Hung 2004).
|Continuing decline in area, extent and/or quality of habitat:||Yes|
|Generation Length (years):||16-17|
|Movement patterns:||Not a Migrant|
|Use and Trade:||There does not appear to be trade (for consumption) in most areas of China and Japan. However, Narrow-ridged Finless Porpoises are sold and consumed fairly widely in certain fishing ports/markets in South Korea (e.g., Busan, Ulsan). A limited number of Narrow-ridged Finless Porpoises are sold by fishermen to local display/entertainment institutions.|
Finless Porpoises, like other phocoenids (Jefferson and Curry 1994), are extremely susceptible to entanglement in gillnets, and large numbers have been, and continue to be, killed in many parts of their range.
In Japan, Narrow-ridged Finless Porpoises become entangled in a variety of types of fishing gear (Amano 2009). Changes in fishing methods and the use of acoustic deterrents may have reduced the incidental catch in some areas such as western Kyushu (Kasuya 1999, Amano et al. 2017), but substantial numbers are still being taken in gillnets and other fishing gear. A total of 114 specimens were collected in Japan during 1985–1992 from western and north-eastern Kyushu including parts of the western Inland Sea (Shirakihara et al. 1993): 84 (73%) of them had been killed incidentally in fisheries, 25 had been found dead on the beach or in the sea, and there was no information on the other five. Fishing gears that killed the 84 porpoises were bottom-set gillnets (58), surface gillnets (17), trap nets (7), trawl nets (1) and drifting (ghost) nets (1). Such fishing gears are common in Japan and probably kill Finless Porpoises off other coasts, although usually such catches go unreported. Shirakihara and Shirakihara (2013) estimated 238-270 Finless Porpoises are bycaught annually in Ariake Sound and Tachibana Bay.
In Korea (information is available only from the Republic of Korea), an official reporting system implemented in 2011 resulted in a count of 2,107 Finless Porpoises bycaught inshore, mainly along the west coast, in 2012 (Kim et al. 2013). Although reporting apparently has improved, there is still assumed to be substantial under-reporting for the following reasons according to Park et al. (2015): “First, each Finless Porpoise caught in a fishing net costs approximately $100 (100,000 Wons) [sic]. Therefore, the dead porpoise can be discarded because it has no significant value. Second, the bycatch of Finless Porpoises is usually reported only in large harbours, with deaths in smaller ports seldom reported (Kim et al., 2013). Therefore, it is likely that the impact of the bycatch of Finless Porpoises on its population is underestimated.”
Yang et al. (1999) reported that Finless Porpoises were the most frequently captured cetaceans in fishing gear along the Chinese coast and estimated that more than 2,000 were taken in 1994, mainly in trawl, gill, and stow nets. In the waters of western Taiwan, including the Matsu and Chinmen Islands (western Taiwan Strait), a considerable number of Finless Porpoises are taken in trammel nets, trawl nets, stow nets, and other gear (J. Y. Wang unpublished data). Given the numbers and types of net fisheries in Chinese coastal waters (e.g., Zhou and Wang 1994, Yang et al. 1999), there are serious concerns about the level of bycatch of both Finless Porpoise species. Illegal electric fishing became widespread in the Yangtze River during the 1990s, and it may kill porpoises outright and certainly contributes to the depletion of their prey (Reeves et al. 2000).
In some parts of their range, there is an aversion to eating Finless Porpoises (Kasuya 1999, J. Y. Wang unpublished data). However, there is a long history of porpoises taken incidentally being sold for human consumption in at least some parts of Japan (e.g., Mizue et al. 1965) and on the Korean peninsula (IWC 2000). According to M. Amano (pers. comm.), consumption of Finless Porpoises no longer occurs in Japan. An analysis of the species composition of odontocete products in Korean markets during 2003–2004 estimated that the true catch of Finless Porpoises in Korean waters during this period was probably about four times the officially reported catch of 142 animals (Baker et al. 2006).
A survey of levels and drivers of human-caused mortality carried out in 27 fishing settlements along the middle-lower Yangtze channel identified three main categories of observed porpoise deaths: (1) interaction with fishing gear, (2) vessel strikes, and (3) unknown cause (Turvey et al. 2013). A total of 344 dated porpoise mortality events directly observed by informants between 1950 and 2008 were reported, including 75 porpoise deaths from 1989–1998 and 147 from 1999–2008. This apparent increase was said to be driven by the greater number of deaths attributed to vessel strikes (19 to 35) and unknown causes (31 to 94) in the more recent decade. Comparison of the reported mortality with the estimated abundance suggests that the fraction of individuals removed from the declining porpoise population in the Yangtze had quadrupled over the course of two decades. Turvey et al. concluded that entanglement in rolling hook long-lines was probably responsible for the greatest number of porpoise deaths in fishing gear and that vessel strikes and other anthropogenic factors, such as electrofishing, had become the dominant and increasing causes of mortality in the Yangtze.The causes of an apparent decline in Finless Porpoise numbers in the Seto Inland Sea of Japan are not fully understood, but include incidental mortality in fisheries as well as various forms of habitat degradation (IWC 2000, 2006). Parts of the coastal regions of the Inland Sea are highly developed and industrialized, so the associated threats of habitat loss (e.g., from reclamation and sand mining) and pollution are suspected to have had some (undetermined) level of impact (Kasuya et al. 2002).
|Conservation Actions:||Neophocaena phocaenoides (meant at the time of listing to also include N. asiaeorientalis) is listed in Appendix I of the Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES), and in Appendix II of the Convention on Migratory Species (CMS). Management measures are needed to reduce the threats, particularly incidental mortality in fisheries and vessel strikes, and also to protect (or if possible restore) the environmental conditions needed for these animals to survive and reproduce.|
Amano, M. 2002. Finless porpoise Neophocaena phocaenoides. In: W. F. Perrin, B. Wursig and J. G. M. Thewissen (eds), Encyclopedia of Marine Mammals, pp. 432-435. Academic Press.
Amano, M. 2009. Finless porpoise Neophocaena phocaenoides. In: W. F. Perrin, B. Würsig and J. G. M. Thewissen (eds) (eds), Encyclopedia of Marine Mammals, pp. 437-439. Elsevier, Amsterdam.
Amano, M., Kusumoto, M., Abe, M. and Akamatsu, T. 2017. Long-term effectiveness of pingers on a small population of finless porpoises in Japan. Endangered Species Research 32: 35-40.
Amano, M., Miyazaki, N. and Kureha, K. 1992. A morphological comparison of skulls of the finless porpoise Neophocaena phocaenoides from the Indian Ocean, Yangtze River and Japanese waters. Journal of the Mammalogical Society of Japan 17: 59-69.
Amano, M., Nakahara, F., Hayano, A. and Shirakihara, K. 2003. Abundance estimate of finless porpoises off the Pacific coast of eastern Japan based on aerial surveys. Mammal Study 28: 103-110.
Baker, C. S., Lukoschek, V., Lavery, S., Dalebout, M., Yong-Un, M., Endo, T. and Funahashi, N. 2006. Incomplete reporting of whale, dolphin and porpoise 'bycatch' revealed by molecular monitoring of Korean markets. Animal Conservation 9: 474-482.
Chen, L. Bruford, M.W., Xu, S, Zhou, K. and Yang G. 2010. Microsatellite variation and significant population genetic structure of endangered finless porpoises (Neophocaena phocaenoides) in Chinese coastal waters and the Yangtze River. Marine Biology 157: 1453-1462.
Chen, L., Bruford, M.W., Xu, S., Zhou, K. and Yang, G. 2010. Microsatellite variation and significant population genetic structure of endangered finless porpoises (Neophocaena phocaenoides) in Chinese coastal waters and the Yangtze River. Marine Biology 157: 1453-1462.
Chen, M., Zheng, J., Wu, M. Ruan, R., Zhao, Q. and Wang, D. 2014. Genetic diversity and population structure of the Critically Endangered Yangtze finless porpoise (Neophocaena asiaeorientalis asiaeorientalis) as revealed by mitochondrial and microsatellite DNA. International Journal of Molecular Sciences 15: 11307-11323.
Gao, A. 1991. Morphological differences and genetic variations among the populations of Neophocaena phocaenoides. Nanjing Normal University.
Gao, A. and Zhou, K. 1995. Geographical variation of external measurements and three subspecies of Neophocaena phocaenoides in Chinese waters. Acta Theriologica Sinica 15: 81-92.
Hashimoto, M. Shirakihara, K. and Shirakihara, M. 2015. Effects of bycatch on the population viability of the narrow-ridged finless porpoises in Ariake Sound and Tachibana Bay, Japan. Endangered Species Research 27: 87-94.
IUCN. 2017. The IUCN Red List of Threatened Species. Version 2017-3. Available at: www.iucnredlist.org. (Accessed: 5 December 2017).
Iwata, H., Tanabe, S., Miyazaki, N. and Tatsukawa, R. 1994. Detection of butyltin compound residues in the blubber of marine mammals. Marine Pollution Bulletin 28: 607–612.
Iwata, H., Tanabe, S., Mizuno, T. and Tatsukawa, R. 1995. High accumulation of toxic butyltins in marine mammals from Japanese coastal waters. Environmental Science and Technology 29: 2959-2962.
IWC. 2000. Report of the standing sub-committee on small cetaceans. Journal of Cetacean Research and Management 2 (Supplement), 235–263.
IWC. 2006. Report of the sub-committee on small cetaceans. Journal of Cetacean Research and Management 8: 221-232.
Jefferson, T.A. 2002. Preliminary analysis of geographic variation in cranial morphometrics of the finless porpoise (Neophocaena phocaenoides). Raffles Bulletin of Zoology Supplement 10: 3-14.
Jefferson, T.A., and Curry, B.E. 1994. A global review of porpoise (Cetacea: Phocoenidae) mortality in gillnets. Biological Conservation 67: 167–183.
Jefferson, T. A. and Hung, S. K. 2004. Neophocaena phocaenoides. Mammalian Species 746: 1-12.
Jefferson, T.A. and Wang, J.Y. 2011. Revision of the taxonomy of finless porpoises (genus Neophocaena): the existence of two species. Journal of Marine Animals and Their Ecology 4(1): 3-16.
Jia, K, Lin, W., Gui, D., Karczmarski, L. and Wu, Y. 2014. Molecular evidence reveals the distinctiveness of Indo‑Pacific finless porpoises (Neophocaena phocaenoides) in the Pearl River Estuary and insights into genus Neophocaena’s origin. Marine Biology 161: 1919-1930.
Ju, J. Yang. M., Xu, S., Zhou, K. and Yang, G. 2012. High level population differentiation of finless porpoises (Neophocaena phocaenoides) in Chinese waters revealed by sequence variability of four nuclear introns. Molecular Biology Reports 39: 7755-7762.
Ju, J. Yang. M., Xu, S., Zhou, K. and Yang, G. 2012. High level population differentiation of finless porpoises (Neophocaena phocaenoides) in Chinese waters revealed by sequence variability of four nuclear introns. Molecular Biology Reports 39: 7755-7762.
Kannan, N., Tanabe, S., Ono, M. and Tatsukawa, R. 1989. Critical evaluation of polychlorinated biphenyl toxicity in terrestrial and marine mammals: increasing impact of non-ortho and mono-ortho coplanar polychlorinated biphenyls from land to ocean. Archives of Environmental Contamination and Toxicology 18: 850-857.
Kasuya, T. 1999. Finless porpoise Neophocaena phocaenoides (G. Cuvier, 1829). In: S. H. Ridgway and R. Harrison (eds), Handbook of Marine Mammals, pp. 411-442. Academic Press, London, UK.
Kasuya, T., Yamamoto, Y. and Iwatsuki, T. 2002. Abundance decline in the finless porpoise population in the Inland Sea of Japan. Raffles Bulletin of Zoology Supplement 10: 57-65.
Le, T. H. L., Takahashi, S., Saeki, K., Nakatani, N., Tanabe, S., Miyazaki, N. and Fujise, Y. 1999. High percentage of butyltin residues in total tin in the livers of cetaceans from Japanese coastal waters. Environmental Science and Technology 33: 1781-1786.
Li, L., Jiang, J., Wang, X. and Jiang, X. 2013. Nested clade phylogeographical analysis of the finless porpoise (Neophocaena phocaenoides) inhabiting Chinese and Japanese coasts. Genetics and Molecular Research 12: 2528-2536.
Lin, W., Frère, C.H., Karczmarski, L., Xia, J., Gui, D., and Wu, Y. 2014. Phylogeography of the finless porpoise (genus Neophocaena): testing the stepwise divergence hypothesis in the northwestern Pacific. Scientific Reports 4: 6572.
Li, S., Xu, S, Wan, H, Ki, H, Zhou, K, and Yang, G. 2013. Genome-wide SNP and population divergence of finless porpoises. Genome Biology and Evolution 5: 758-768.
Liu, R., Wang, D. and Zhou, K. 2000. Effects of water development on river cetaceans in China. In: R.R. Reeves, B.D. Smith and T. Kasuya (eds), Biology and conservation of freshwater cetaceans in Asia, pp. 40-42. IUCN Species Survival Commission Occasional Paper No. 23, Gland, Switzerland.
Li, X., Liu, Y., Tzika, A.C., Zhu, Q., Van Doninck, K. and Milinkovitch, M.C. 2011. Analysis of global and local population stratification of finless porpoises Neophocaena phocaenoides in Chinese waters. . Marine Biology 158: 1791-1804.
Mei, Z., Zhang, X., Huang, S.-L., Zhao, X., Hao, Y., Zhang, L., Qian, Z., Zheng, J., Wang, K. and Wang, D. 2014. The Yangtze finless porpoises: on an accelerating path to extinction? Biological Conservation 172: 117–123.
Mizue, K., Yoshida. K. and Masaki, Y. 1965. Studies on the little toothed whales in the west sea area of Kyushu-XII. Neomeris phocaenoides, so-called Japanese "SUNAMERI", caught in the coast of Tachibana Bay, Nagasaki Pref. Bulletin of the Faculty of Fisheries, Nagasaki University 18: 7-29.
Moore J. 2015. Intrinsic growth (rmax) and generation time (T) estimates for odontocetes of the genus Sousa, Orcaella, and Neophocaena, in support of IUCN Red List Assessments. NOAA Technical Memorandum NMFS SWFSC.
Park, K. J., An, D.H., Lim, C. W., Lee, T-H. and Kim, D. N. 2012. A mass mortality of the finless porpoise Neophocaena asiaeorientalis at a dike of the Saemangeum Sea: possible effects of unusually low temperatures. . Korean Journal of Fisheries and Aquatic Sciences 45: 723-729.
Pilleri, G. and Gihr, M. 1972. Contribution to the knowledge of the cetaceans of Pakistan with particular reference to the genera Neomeris, Sousa, Delphinus and Tursiops and description of a new Chinese porpoise (Neomeris asiaeorientalis). Investigations on Cetacea 4: 107-162.
Pilleri, G. and Gihr, M. 1974. Contribution to the knowledge of the cetaceans of southwest and monsoon Asia (Persian Gulf, Indus Delta, Malabar, Andaman Sea and Gulf of Siam). Investigations on Cetacea 5: 95-153.
Reeves, R.R., Jefferson, T.A., Kasuya, T., Smith, B.D., Wang, D., Wang, P., Wells, R.S., Wursig, B. and Zhou, K. 2000. Report of the Workshop to Develop a Conservation Action Plan for the Yangtze River Finless Porpoise, Ocean Park, Hong Kong, 16-18 September 1997. In: R. R. Reeves and B. D. Smith and T. Kasuya (eds), Biology and conservation of freshwater cetaceans in Asia, pp. 67-80. Occasional Paper of the IUCN Species Survival Commission.
Reeves, R.R., Smith, B.D., Crespo, E.A. and di Sciara, G.N. (compilers) 2003. Dolphins, Whales and Porpoises: 2002-2010 Conservation Action Plan for the World's Cetaceans. IUCN/SSC Cetacean Specialist Group. IUCN, Gland, Switzerland and Cambridge, UK.
Reeves, R.R., Wang, J.Y. and Leatherwood, S. 1997. The finless porpoise, Neophocaena phocaenoides (G. Cuvier 1829): a summary of current knowledge and recommendations for conservation action. Asian Marine Biology 14: 111-143.
Rice, D.W. 1998. Marine Mammals of the World: Systematics and Distribution. Allen Press, Lawrence, Kansas, USA.
Shirakihara, K. and Shirakihara, M. 2002. Survey of Ariake/Tachibana Bay, Omura Bay and Inland Sea. Survey of Marine Animals: finless porpoises, pp. 27–52. Biodiversity Centre, Fujiyoshida., Fujiyoshida.
Shirakihara, K., Shirakihara, M. and Yamamoto, Y. 2007. Distribution and abundance of finless porpoise in the Inland Sea of Japan. Marine Biology 150: 1025-1032.
Shirakihara, K., Yoshida. H., Shirakihara, M. and Takemura, A. 1992. A questionnaire survey on the distribution of the finless porpoise, Neophocaena phocaenoides, in Japanese waters. Marine Mammal Science 8: 160-164.
Shirakihara, M., Takemura, A. and Shirakihara, K. 1993. Age, growth, and reproduction of the finless porpoise, Neophocaena phocaenoides, in the coastal waters of western Kyushu, Japan. Marine Mammal Science 9: 392-406.
Smith, B. D., Sinha, R. K., Zhou, K., Chaudhry, A. A., Renjun, L., Wang, D., Ahmed, B., Haque, A. K. M. and Sapkota, K. 2000. Register of water development projects affecting Asian river cetaceans. In: R.R. Reeves, B. D. Smith, and T. Kasuya (eds), Biology and Conservation of Freshwater Cetaceans in Asia, pp. 22–39. IUCN/SSC Occasional Paper No. 23, Gland, Switzerland and Cambridge, UK.
Taylor, B.L., Chivers, S.J., Larese, J. and Perrin, W.F. 2007. Generation length and percent mature estimates for IUCN assessments of Cetaceans. NOAA Southwest Fisheries Science Center, La Jolla, California. Administrative Report LJ-07-01.
Turvey, S.T., Risley, C.L., Moore, J.E., Barrett, L.A., Hao, Y., Zhao, X., Zhou, K. and Wang, D. 2013. Can local ecological knowledge be used to assess status and extinction drivers in a threatened freshwater cetacean? Biological Conservation 157: 352-360.
Wang, D. 2009. Population status, threats and conservation of the Yangtze finless porpoise. Chinese Science Bulletin 54: 3473-3484.
Wang, D., Liu, R., Zhang, X, Yang, J., Wei, Z., Zhao, Q. and Wang, X. 2000. Status and conservation of the Yangtze Finless Porpoise. In: R. R. Reeves, B. D. Smith, and T. Kasuya (eds), Biology and Conservation of Freshwater Cetaceans in Asia, pp. 81–85. IUCN SSC Occasional Paper No. 23, Gland, Switzerland and Cambridge, UK.
Wang, D., Turvey, S.T., Zhao, X. and Mei, Z. 2013. Neophocaena asiaeorientalis ssp. asiaeorientalis. The IUCN Red List of Threatened Species 2013.
Wang, J.Y., Frasier, T.R., Yang, S.C. and White, B.N. 2008. Detecting recent speciation events: the case of the finless porpoise (genus Neophocaena). Heredity 101: 145-155.
Wang, J.Y., Yang, S.C., Wang, B.J. and Wang, L.S. 2010. Distinguishing between two species of finless porpoises (Neophocaena phocaenoides and N. asiaeorientalis) in areas of sympatry. Mammalia 74.
Wang, P. 1992a. On the taxonomy of the finless porpoise in China. Fisheries Science 11(6): 10-14.
Wang, P. 1992b. The morphological characters and the problem of subspecies identifications of the finless porpoise. Fisheries Science 11: 4-9.
Xu, S., Ren, W., Zhou, X., Zhou, K. and Yang, G. 2010. Sequence polymorphism and geographical variation at a positively selected MHC-DRB gene in the finless porpoise (Neophocaena phocaenoides): implication for recent differentiation of the Yangtze finless porpoise? Journal of Molecular Evolution 71: 6-22.
Yang, G., Guo, L., Bruford, M.W., Wei, F. and Zhou, K. 2008. Mitochondrial phylogeography and population history of finless porpoises in Sino-Japanese waters. Biological Journal of the Linnaean Society 95: 193-204.
Yang, G., Zhou, K.,Gao, A. and Ching, Q. 1998. A study on the life table and dynamics of three finless porpoise populations in the Chinese waters. Acta Theriologica Sinica 18: 1-7.
Yang, G., Zhou, K. Y., Xu, X. R. and Leatherwood, S. 1999. A survey on the incidental catches of small cetaceans in coastal waters of China. Chinese Journal of Applied Ecology 10(6): 713-716.
Yoshida, H. 2002. Population structure of finless porpoises (Neophocaena phocaenoides) in coastal waters of Japan. Raffles Bulletin of Zoology Supplement 10: 35-42.
Yoshida, H., Higashi, N., Ono, H. and Uchida, S. 2010. Finless porpoise (Neophocaena phocaenoides) discovered at Okinawa Island, Japan, with the source population inferred from mitochondrial DNA. Aquatic Mammals 36: 279-283.
Yoshida, H., Shirakihara, K., Shirakihara, M. and Takemura, A. 1995. Geographic variation in the skull morphology of the finless porpoise Neophocaena phocaenoides in Japanese waters. Fisheries Science 61: 555-558.
Yoshida, H., Yoshioka, M., Shirakihara, M. and Chow, S. 2001. Population structure of finless porpoises Neophocaena phocaenoides) in coastal waters of Japan based on mitochondrial DNA sequences. Journal of Mammalogy 81: 123-130.
Zhang, C. I., Park, K. J., Kim, Z. G. and Sohn, H. 2005. Abundance estimation of finless porpoise Neophocaena phocaenoides from ship surveys in the west coast of Korea. IWC Scientific Committee Document SC/57/SM15. Available from IWC Secretariat. Cambridge, UK.
Zhang, X., Liu Renjun, Zhao Qingshong, Zhang Guocheng, Sei Zhuo, Wang Xiaoqiang and Yang Jian. 1993. The population of finless porpoise in the middle and lower reaches of Yangtze River. Acta Theriologica Sinica 13: 260-270.
Zhao, X., Barlow, J., Taylor, B.L., Pitman, R.L., Wang, K., Wei, W., Stewart, B.S., Turvey, S.T., Akamatsu, T., Reeves, R.R. and Wang, D. 2008. Abundance and conservation status of the Yangtze finless porpoise in the Yangtze River, China. Biological Conservation 141: 3006-3018.
Zheng, J.-S, Xia, J.-H,. He, S.-P. and Wang, D. 2005. Population genetic structure of the Yangtze finless porpoise (Neophocaena phocaenoides asiaeorientalis): implications for management and conservation. Journal of Molecular Evolution 71: 6-22.
Zhou, K. and Wang, X. 1994. Brief review of passive fishing gear and incidental catches of small cetaceans in Chinese waters. In: Perrin, W.F., Donovan, G.,P. and Barlow, J. (eds), Gillnets and Cetaceans Report of the International Whaling Commission 15, pp. 347-354. Cambridge, U.K.
Zhou, K., Yang, G., Gao, A., Sun, J. and Xu, X. 2000. Abundance and distribution of finless porpoises in the Nanjing-Hukou section of the lower Yangtze River. In: R. R. Reeves, B. D. Smith and T. Kasuya (eds), Biology and conservation of freshwater cetaceans in Asia, pp. 91-96. Occasional Paper of the IUCN Species Survival Commission.
|Citation:||Wang, J.Y. , Reeves, R. 2017. Neophocaena asiaeorientalis. The IUCN Red List of Threatened Species 2017: e.T41754A50381766.Downloaded on 17 July 2018.|
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