Bibliography [top]

Bannikov, A. G. 1981. Khulan. Lesnaya Promiyshlennosti, Moscow, Russia.

Batsaikhan, N., Buuveibaatar, B., Chimed, B., Enkhtuya, O., Galbrakh, D., Ganbaatar, O., Lkhagvasuren, B., Nandintsetseg, D., Berger, J., Calabrese, J.M., Edwards, A.E., Fagan, W.F., Fuller, T.K., Heiner, M., Ito, T.Y., Kaczensky, P., Leimgruber, P., Lushchekina, A., Milner-Gulland, E.J., Mueller, T., Murray, M.G., Olson, K.A., Reading, R., Schaller, G.B., Stubbe, A., Stubbe, M., Walzer, C., von Wehrden, H., Whitten, T. 2014. Conserving the World's Finest Grassland Amidst Ambitious National Development. Conservation Biology 28(6): 1736–1739.

Bennett, E.A., Champlot, S., Pruvost, M., Gautier, M., Arbuckle, B., Balasescu, A., Eisenmann, V., Kaczensky, P., Kuehn, R., Mashkour, M., Morales Muniz, A., Peters, J., Tournepiche, J.-F., Uerpmann, H.-P., Walzer, C., Grange, T., Geigl, E.-M. 2012. The Decline of the Asiatic Wild Ass Recording 100,000 Years of Genetic Diversity of the Species. Page 26 in: P. Kaczensky and J. Ransom (Eds). International Wild Equid Conference - Book of abstracts. Conference in Vienna, Austria from 12-22 September 2012. Research Institute of Wildlife Ecology, University of Veterinary Medicine, Austria.

Bi, J.H. 2007. A Study on the Status of Asiatic Wild Ass (Equus hemionus hemionus) and its Ecological Problems. PhD thesis 135 at the Beijing, Beijing Forestry University. [in Chinese].

Bull, J. W., Suttle, K. B., Singh, N. J., and Milner-Gulland, E. J. 2013. Conservation when nothing stands still: moving targets and biodiversity offsets. Frontiers in Ecology and the Environment 11: 203-210.

Business and Biodiversity Offsets Programme (BBOP). 2012. Standard on Biodiversity Offsets. BBOP, Washington, D.C..

Chu, H. 2008. Diets, Populations and Habitats of Khulan (E. hemionus) and Goitred Gazelle (G. subgutturosa) in the Mt. Kalamaili Ungulate Nature Reserve, Xinjiang, China. PhD thesis 183 at the Chinese Academy of Sciences. [in Chinese].

Crees, J.J., and Turvey, S.T. 2014. Holocene extinction dynamics of Equus hydruntinus, a late-surviving European megafaunal mammal. Quaternary Science Reviews 91: 16-29.

Denzau, G., and Denzau, H. 1999. Wildesel. Jan Thorbecke Verlag, Stuttgart, Germany. [in German].

Feh, C., Boldsukh, T. and Tourenq, C. 1994. Are family groups in equids a response to cooperative hunting by predators? The case of Mongolian kulans (Equus hemionus luteus Matschie). Revue d'Écologie (La Terre et la Vie) 49: 11-20.

Feh, C., Munkhtuya, B., Enkhbold, S. and Sukhbaatar, T. 2001. Ecology and Social Structure of the Gobi Khulan (Equus hemionus subsp). in the Gobi B National Park, Mongolia. Biological Conservation 101: 51-61.

Feh, C., Shah, N., Rowen, M., Reading, R. and Goyal, S. P. 2002. Status and action plan for the Asiatic wild ass (Equus hemionus). In: P. D. Moehlman (ed.), Equids: Zebras, Asses and Horses. Status Survey and Conservation Action Plan, pp. 62-71. IUCN, Gland, Switzerland.

Gee, E. P. 1963. The Indian wild ass: a survey. Journal of the Bombay Natural History Society 60: 517-529.

Goyal, S. P., Sinha, B., Shah, N. and Panwar, H. S. 1999. Sardar Sarovar Project – a conservation threat to the Indian wild ass (Equus hemionus khur). Biological Conservation 88: 277-284.

Grubb, P. 2005. Artiodactyla. In: D.E. Wilson and D.M. Reeder (eds), Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), pp. 637-722. Johns Hopkins University Press, Baltimore, USA.

Grubb, P. 2005. Order Perissodactyla. In: D. E. Wilson and D. M. Reeder (eds), Mammal Species of the World, pp. 629-636. The Johns Hopkins University Press, Baltimore, Maryland, USA.

Gueta, T., Templeton, A.R., Bar-David, S. 2014. Development of genetic structure in a heterogeneous landscape over a short time frame: the reintroduced Asiatic wild ass. Conservation Genetics 15: 1231-1242.

Hamidi, A. h. K., Jowkar, H., and Nabiyan, M. 2012. Anthropogenic Threats to Persian Onager (Equus hemionus onager) in Iran. Page 53 in: P. Kaczensky and J. Ransom (Eds). International Wild Equid Conference - Book of abstracts. Conference in Vienna, Austria from 12-22 September 2012. Research Institute of Wildlife Ecology, University of Veterinary Medicine, Austria..

Hemami, M.-R. and Momeni, M. 2013. Estimating abundance of the Endangered onager Equus hemionus onager in Qatruiyeh National Park, Iran. Oryx 47: 266-272.

Heptner, V. G., Nasimovich, A. A., and Bannikov, A. G. 1988. Kulan: Equus (Equus) hemionus. Pages 1011-1036 in: Mammals of the Soviet Union Volume 1 – Artiodactyla and Perissodactyla. [English translation of the original book published in 1961 by Vysshaya Shkola Publishers Moscow]. Smithsonian Institution Libraries and The National Science Foundation Washington, D.C., USA..

IUCN. 2015. The IUCN Red List of Threatened Species. Version 2015-4. Available at: (Accessed: 19 November 2015).

Kaczensky, P. and Linnell, J.D.C. 2015. Rapid assessment of the mammalian community of the Badhyz Ecosystem, Turkmenistan, October 2014. Norwegian Institute for Nature Research, NINA Report 1148..

Kaczensky, P., Dresley, V., Vetter, D., Otgonbayar, H., Walzer, C. 2010. Water use of Asiatic wild asses in the Mongolian Gobi. Exploration into the Biological Resources of Mongolia (Halle/Saale, Germany) 11: 291-298.

Kaczensky, P., Ganbaatar, O., von Wehrden, H., Walzer, C. 2008. Resource selection by sympatric wild equids in the Mongolian Gobi. Journal of Applied Ecology 45: 1762–1769.

Kaczensky, P., Ganbataar, O., Altansukh, N., Enkhsaikhan, N., Stauffer, C., Walzer, C. 2011. The Danger of Having All your Eggs in One Basket – Winter Crash of the Re-introduced Przewalski’s Horses in the Mongolian Gobi. PloS ONE 6(12): e28057, doi:10.1371/journal.pone.0028057.

Kaczensky, P., Kuehn, R., Lhagvasuren, B., Pietsch, S., Yang, W. and Walzer, C. 2011. Connectivity of the Asiatic wild ass population in the Mongolian Gobi. Biological Conservation 144: 920-929.

Kaczensky, P., Sheehy, D. P., Johnson, D. E., Walzer, C., Lhkagvasuren, D. and Sheehy, C. M. 2006. Room to roam? The threat to khulan (wild ass) from human intrusion. Mongolia Discussion Papers, East Asia and Pacific Environment and Social Development Departure, World Bank: 69.

Karlstetter, M., Mallon, D. 2014. Assessment of gaps and needs in migratory mammal conservation in Central Asia. Report prepared for the Convention on the Conservation of Migratory Species of Wild Animals (CMS) and the Deutsche Gesellschaft für Internationale Zusammenarbeit (GIZ) GmbH. Financed by the Ecosystem Restoration in Central Asia (ERCA) component of the European Union Forest and Biodiversity Governance Including Environmental Monitoring Project (FLERMONECA).

Klingel, H. 1998. Observations on social organization and behavior of African and Asiatic wild asses (Equus africanus and E. hemionus). Applied Animal Behaviour Science 60: 103-113.

Kuznetsov, V. 2014. Return of Onager. International Journal Turkmenistan 1-2: 84-92 [in Russian and English].

Levanov, V.F., Sokolov, S.V., Kaczensky, P. 2013. Corral mass capture device for Asiatic wild asses Equus hemionus. Wildlife Biology 19: 325-334.

Lkhagvasuren, B. 2007. Population assessment of khulan (Equus hemionus) in Mongolia. Erforschung Biologischer Ressourcen der Mongolei 10: 45-48.

Lukarevski, V. S., and Gorelov, Y. K. 2007. Khulan (Equus hemionus Pallas 1775) in Turkmenistan. Exploration into the Biological Resources of Mongolia (Halle/Saale) 10: 231-240.

Meldebekov, A. M., M. K. Bajzhanov,Bekenov, A. B., and Kovshar, A. F. 2010. The Red Data Book of the Republic of Kazakhstan, Volume 1: Animals, part 1: Vertebrates. 4th Edition. Almaty, Kazakstan.

Neumann-Denzau, G. and Denzau, H. 2007. Remarks on the social system of the Mongolian wild ass (Equus hemionus hemionus). Exploration into the Biological Resources of Mongolia (Halle/Saale) 10: 177-187.

Norton-Griffiths, M., Frederick, H., Slaymaker, D. M., and Payne, J. 2013. Preliminary estimates of wildlife and livestock populations in the Oyu Tolgoi Area of the south-eastern Gobi Desert, Mongolia, May - July 2013. Unpublished preliminary report to Oyu Tolgoi: 11pp.

Norton-Griffiths, M., Frederick, H., Slaymaker, D.M., Payne, J. 2015. Aerial Census of Wildlife and Livestock in the Oyu Tolgoi Area of the Gobi Desert, Mongolia May - July 2013. Final Report to Oyu Tolgoi LLC. 200 pp. .

Nowzari, H., Hemami, M.-R., Karami, M., Zarkesh, M.M.K., Riazi, B., Rubenstein, D.I. 2012. Population Parameters of Persian Wild Ass (Equus hemionus onager) in Qatrouyeh National Park, Iran. Page 85 in: P. Kaczensky and J. Ransom (Eds). International Wild Equid Conference - Book of abstracts. Conference in Vienna, Austria from 12-22 September 2012. Research Institute of Wildlife Ecology, University of Veterinary Medicine, Austria.

Oakenfull, E.A. and Ryder, O.A. 2002. Genetics of equid species and subspecies. In: P.D. Moehlman (ed.), Equids: Zebras, Asses and Horses, pp. 108-112. IUCN Publication Services Unit, Cambridge, United Kingdom.

Oakenfull, E.A., Lim, H.N. and Ryder, O.A. 2000. A survey of equid mitochondrial DNA: implications for the evolution, genetic diversity and conservation of Equus. Conservation Genetics 1: 341-255.

Pavlov, M. P. 1996. Translocations of kulans in the former Soviet Union. Newsletter of the Re-introduction specialist group of IUCN' Species Survival Commission (SSC) 12: 15-16.

Plakhov, K. N., Sokolov, S. V., Levanov, V. F., and Akylbekova, A. Z. 2012. News in Kulan reintroduction in Kazakhstan. In: Meldebekov, A.M., Bekenov, A. B. Grachev, Yu. A. Baydavletov, R. Zh. Sklyarenko, S. L. Bodrova, N. P. (Eds.); Zoological and game management researches in Kazakhstan and adjacent countries. Materials on international theoretical and practical conference devoted to centenary of birth of the founder of Kazakhstan’s theriology and game management schools, a Laureate of State Prizes of USSR and KazSSR, a Corresponding Member of AS of KazSSR – Arkadiy Alexandrovich Sludskiy (Almaty, 1-2 March 2012), Almaty, Kazakhstan, 151-153 [In Russian].

Ransom, J.I., Kaczensky, P., Lubow, B.C., Ganbaatar, O., Altansukh, N. 2012. A collaborative approach for estimating terrestrial wildlife abundance. Biological Conservation. Biological Conservation 153: 219–226.

Reading, R. P., Mix, H., Lhagvasuren, B., Feh, C., Kane, D. P., Dulamtseren, S. and Enkhbold, S. 2001. Status and distribution of khulan (Equus hemionus) in Mongolia. Journal of the Zoology 254: 381-389.

Renan, S., Greenbaum, G., Shahar, N., Templeton, A.R., Bouskila, A., Bar-David, S. 2015. Stochastic modelling of shifts in allele frequencies reveals a strongly polygynous mating system in the re-introduced Asiatic wild ass. Molecular Ecology 24: 1433-1446.

Rosenbom, S., Costa, V., Chen, S., Khalatbari, L., Yusefi, G.H., Abdukadir, A., Yangzom, C., Kebede, F., Teclai, R., Yohannes, H., Hagos, F., Moehlman, P.D. and Beja-Pereira, A. 2015. Reassessing the evolutionary history of ass-like equids: insights from patterns of genetic variation in contemporary extant populations. Molecular Phylogenetic Evolution 85: 88-96.

Rubenstein, D. I. 1986. Life history and social organization in arid adapted ungulates. In: D. I. Rubenstein and R. W. Wrangham (eds), Ecological Aspects of social evolution, pp. 282-302. Princeton University Press, Princeton.

Rubenstein, D. I., Sundaresan, S., Fischhoff and Saltz, D. 2007. Social Networks in Wild Asses: Comparing Patterns and Processes among Populations. Exploration into the Biological Resources of Mongolia 10: 159-176.

Saltz, D. and Rubenstein, D. I. 1995. Population dynamics of a reintroduced Asiatic wild ass (Equus hemionus) herd. Ecological Applications 5: 327-335.

Saltz, D., Rubenstein, D.I., White, G.C. 2006. The impact of increased environmental stochasticity due to climate change on the dynamics of Asiatic wild ass. Conservation Biology 20: 1402-1409.

Shah, N. V. 1993. Ecology of wild ass (Equus hemionus khur) in Little Rann of Kutch. Ph.D. Thesis, M.S. University of Baroda.

Shah, N. V. and Qureshi, Q. 2007. Social organization and determinants of spatial distribution of khur (Equus hemionus khur). Exploration into the Biological Resources of Mongolia 10: 189-200.

Singh, N.J., Milner-Gulland, E.J. 2011. Conserving a moving target: planning protection for a migratory species as its distribution changes. Journal of Applied Ecology 48: 35-46.

Solomatin, A. O. 1973. The Wild Ass. Nauka, Moscow [in Russian].

Stubbe, A., Stubbe, M. and Batsajchan. 2007. Morphology, reproduction and mortality of Equus hemionus hemionus in Mongolia. Erforschung Biologischer Ressourcen der Mongolei (Halle/Saale) 2007(10): 117-132.

Stubbe, A., Stubbe, M., Batsaikhan, N., Samjaa, R. 2012. Long-term ecology of Asiatic wild ass (Equus h. hemionus Pallas) in Central Asia. Exploration into the Biological Resources of Mongolia (Halle/Saale, Germany) 12: 61-76.

Stubbe, A., Stubbe, M., Batsajchan, N., Samjaa, R. and Doržderem, S. 2005. First results of wild ass research in the South Gobi Aymag, Mongolia in 2003 and 2004. Erforschung Biologischer Ressourcen der Mongolei (Halle/Saale) 9: 107-120.

Sundaresan, S.R., Fischhoff, I.R., Dushoff, J., Rubenstein, D.I. 2007. Network metrics reveal differences in social organization between two fission-fusion species, Grevy's zebra and onager. Oecologia 151: 140-149.

Tatin, L., Darreh-Shoori, B., Tourenq, C., Tatin, D., Azmayesh, B., Zeh-Zad, B. and Rezahi. 2003. The last populations of the critically endangered onager Equus hemionus onager in Iran: urgent requirements for protection and study. Oryx 37: 488-491.

Volf, J. 2010. Sixty years of kulan, Equus hemionus kulan (Groves et Mazák, 1967) breeding at Prague Zoo. Equus: 30-55.

Von Wehrden, H., Hanspach, J., Kaczensky, P., Fischer, J., Wesche, K. 2012. A global assessment of the non-equilibrium concept in rangelands. Ecological Applications 22: 393–399.

Warner, S. 2014. Road-kills of Asiatic Wild Ass in the Central Negev – possible causes and means to prevent them. Unpublished report, Hebrew University of Jerusalem, Israel, 11pp. [in Hebrew].

Wingard, J.R. and Zahler, P. 2006. Silent Steppe: The Illegal Wildlife Trade Crisis in Mongolia. Mongolia Discussion Papers, East Asia and Pacifi c Environment and Social Development Department. Washington D.C.: World Bank.

Xia, C., Cao, J., Zhang, H., Gao, X., Yang, W., Blank, D. 2014. Reintroduction of Przewalski’s horse (Equus ferus przewalskii) in Xinjiang, China: The status and experience. Biological Conservation 177: 142-147.

Xu, W., Xia, C., Yang, W., Blank, D.A., Qiao, J., Liu, W. 2012. Seasonal diet of Khulan (Equidae) in Northern Xinjiang, China. Italian Journal of Zoology 79: 92-99.

Citation: Kaczensky, P., Lkhagvasuren, B., Pereladova, O., Hemami, M. & Bouskila, A. 2015. Equus hemionus. The IUCN Red List of Threatened Species 2015: e.T7951A45171204. . Downloaded on 29 November 2015.
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