Ursus arctos 

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Abbas, F., Bhatti, Z.I., Haider, J., and Mian, A. 2015. Bears in Pakistan: distribution, population biology and human conflicts. Journal of Bioresource Management 2: 1-13.

Aryal, A., Sathyakumar, S., and Schwartz, C.C. 2010. Current status of brown bears in the Manasalu Conservation Area, Nepal. Ursus 21: 109-114.

Bellemain, E., Swenson, J. E., Tallmon, D., Taberlet, P. and Brunberg, S. 2005. Estimating population size of elusive animals with DNA from hunter-collected feces: Four methods for brown bears. Conservation Biology 19: 150-161.

Brown, D. E. 1985. The grizzly in the southwest. Documentary of an extinction. University of Oklahoma Press, Norman, Oklahoma, USA.

Burgess, E.A., Stoner, S.S., and Foley, K.E. 2014. Brought to bear” an analysis of seizures across Asia (2000-2001). TRAFFIC, Petaling Jaya, Selangor, Malayasia.

Can, O. E. and Togan, I. 2004. Status and management of brown bears in Turkey. Ursus 15(1): 48-53.

Chapron, G., Kaczensky, P., Linnell, J.D.C., von Arx, M., Huber, D., Andrén, H., Vicente López-Bao, J., Adamec, M., Álvares, F., Anders, O., Balčiauskas, L., Balys, V., Bedő, P., Bego, F., Blanco, J. C., Breitenmoser, U., Brøseth, H., Bufka, L., Bunikyte, R., Ciucci, P., Dutsov, A., Engleder, T., Fuxjäger, C., Groff, C., Holmala, K., Hoxha, B., Iliopoulos, Y., Ionescu, O., Jeremić, J., Jerina, K., Kluth, Knauer, F., Kojola, I., Kos, I., Krofel, M., Kubala, J., Kunovac, S., Kusak, J., Kutal, M., Liberg, O., Majić, A., Männil, P., Manz, R., Marboutin, E., Marucco, F., Melovski, D., Mersini, K., Mertzanis, Y., Mysłajek, R.W., Nowak, S., Odden, J., Ozolins, J., Palomero, G., Paunović, M., Persson, J., Potočnik, H., Quenette, P.-Y., Rauer, G., Reinhardt, I., Rigg, R., Ryser, A., Salvatori, V., Skrbinšek, T., Stojanov, A., Swenson, J.E., Szemethy, L., Trajçe, A., Tsingarska-Sedefcheva, E., Váňa, M., Veeroja, R., Wabakken, P., Wölfl, M., Wölfl, S., Zimmermann, F., Zlatanova, D., and Boitani, L. 2014. Recovery of Large carnivores in Europe's modern human-dominated landscapes. Science 346: 1517-1519.

Clark, J. D., Huber, D. and Servheen, C. 2002. Reintroducing bears: lessons and challenges. Ursus 13: 153-163.

Frković, A., Huber, D. and Kusak, J. 2001. Brown bear litter sizes in Croatia. Ursus 12: 103-106.

Gärdenfors, U., Hilton-Taylor, C., Mace, G., and Rodríguez, J.P. 2001. The application of IUCN Red List Criteria at Regional levels. Conservation Biology 15: 1206-1212.

Garshelis, D.L., Gibeau, M.L. and Herrero, S. 2005. Grizzly bear demographics in and around Banff National Park and Kananaskis Country, Alberta. Journal of Wildlife Management 69: 277-297.

Garshelis, D.L., Steinmetz, R., Hajjar, I., and Tabbaa, D. 2015. Brown bear(s) do exist in Syria: fuzzy photo generates much interest. International Bear News 24: 5-8.

Gong, J. and Harris, R.B. 2006. The status of bears in China. Understanding Asian bears to secure their future, pp. 50-56. Japan Bear Network, Ibaraki, Japan.

Guilday, J.E. 1968. Grizzly bears from eastern North America. The American Midland Naturalist 79: 247-250.

Gurung, M. K. 2004. Brown bear observation in Damodar Kunda Valley, Mustang District, Nepal. International Bear News 13(4): 12-14.

Hamdine, W., Thevenot, M. and Michaux, J. 1998. Recent history of the brown bear in the Maghreb. Comptes Rendus de l'Academie des Sciences - Serie III: Sciences de la Vie 321: 565-570.

Haroldson, M. A., F. T. van Manen, and D. D. Bjornlie. 2013. Estimating number of females with cubs-of-the-year. In: F. T. van Manen, M. A. Haroldson, and K. West (eds), Yellowstone grizzly bear investigations: annual report of the Interagency Grizzly Bear Study Team, 2012. U.S., pp. 11–18. Geological Survey, Bozeman, Montana, USA.

Herrero, S. 1994. The Canadian national parks and grizzly bear ecosystems: the need for interagency management. International Conference on Bear Research and Management 9: 7-21.

Hilderbrand, G. V., Jacoby, M. E., Schwartz, C. C., Arthur, S. M., Robbins, C. T., Hanley, T. A. and Servheen, C. 1999. The importance of meat, particularly salmon, to body size, population productivity, and conservation of North American brown bears. Canadian Journal of Zoology 77: 132-138.

IUCN. 2012. Guidelines for Application of IUCN Red List Criteria at Regional and National Levels: Version 4.0. IUCN, Gland, Switzerland and Cambridge, UK.

IUCN. 2017. The IUCN Red List of Threatened Species. Version 2017-1. Available at: (Accessed: 27 April 2017).

IUCN. 2017. The IUCN Red List of Threatened Species. Version 2017-3. Available at: (Accessed: 7 December 2017).

Kaczensky P., G. Chapron, G., von Arx, M., Huber, D., Andrén, C.H., and Linnell, J. 2013. Status, management and distribution of large carnivores – bear, lynx, wolf and wolverine in Europe. Instituto di Ecologia Applicata and with contributions of the IUCN/SSC Large Carnivore Initiative for Europe.

Kasworm, W.F., Proctor, M.F., Servheen, C., and Paetkau, D. 2007. Success of grizzly bear population augmentation in northwest Montana. Journal of Wildlife Management 71: 1261–1266.

Kendall, K.C., Macleod, A.C., Boyd, K.L., Boulanger, J., Royle, J.A., Kasworm, W.F., Paetkau, D., Proctor, M.F., Annis, K., and Graves, T.A. 2016. Density, distribution, and genetic structure of grizzly bears in the Cabinet-Yaak Ecosystem. Journal of Wildlife Management 80: 314-331.

Knott. E.J., Bunnefeld, N.,Huber, D., Reljić, S., Kereži, V., and Milner-Gulland, E.J. 2013. The potential impacts of changes in bear hunting policy for hunting organisations in Croatia. European Journal of Wildllife Research.

Kusak, J. and Huber, D. 1998. Brown bear habitat quality in Gorski kotar, Croatia. Ursus 10: 281-291.

Lamb, C.T., Mowat, G., McLellan, B.M., Nielsen, S.E., and Boutin, S. Submitted. Forbidden fruit: Human settlement and abundant fruit create an ecological trap for an apex omnivore. Journal of Animal Ecology .

Loring, S., and Spiess, A. 2007. Further documentation supporting the former existence of grizzly bears (Ursus arctos) in Northern Quebec-Labrador. Arctic 60: 7-16.

Mace, R.D., Carney, D.W., Chilton-Radant, T., Courville, S.A., Haroldson, M.A., Harris, R.B., Jonkel, J., McLellan, B.N., Madel, M., Manley, R.L., Schwartz, C.C., Servheen, C., Stenhouse, G., Waller, J.S., and Wenum, E. 2012. Grizzly bear population vital rates and trend in the Northern Continental Divide Ecosystem, Montana. Journal of Wildlife Management 76: 119–128.

Manlius, N. 1998. L’Ours brun en Égypte [The brown bear of Egypt]. Écologie 29: 565-581.

Mano, T. 2006. The status of brown bears in Japan. Understanding Asian bears to secure their future, pp. 111-121. Japan Bear Network, Ibaraki, Japan.

Mattson, D. J. 1998. Diet and morphology of extant and recently extinct northern bears. Ursus 10: 479–96.

Mattson, D. J. and Merrill, T. 2002. Extirpations of grizzly bears in the contiguous United States, 1850-2000. Conservation Biology 16: 1123-1136.

McLellan, B. N. 1994. Density dependent population regulation of brown bears. International Conference on Bear Research and Management Monograph Series 3: 15-24.

McLellan, B.N. 2011. 2011. Implications of a high-energy and low-protein diet on body composition, fitness, and competitive abilities of black (Ursus americanus) and grizzly (Ursus arctos) bears. Canadian Journal of Zoology 89: 546-558.

McLellan, B.N. 2015. Some mechanisms underlying variation in vital rates of grizzly bears on a multiple use landscape. Journal of Wildlife Management 79: 749-765.

McLellan, B.N., Hovey, F.W., Mace, R.D., Woods, J.G., Carney, D.W., Gibeau, M.L., Wakkinen, W.L., and Kasworm, W.F. 1999. Rates and causes of grizzly bear mortality in the interior mountains of British Columbia, Alberta, Montana, Washington, and Idaho. Journal of Wildlife Management 63: 911-920.

McLellan, B.N., Mowat, G., Hamilton, A.N., and Hatter, I.W. Submitted. Is the grizzly bear hunt in British Columbia sustainable? Journal of Wildlife Management.

Miller, S.D., Schoen, J.W., Faro, J., and Klein, D.R. 2011. Trends in intensive management of Alaska’s grizzly bears, 1980-2010. Journal of Wildlife Management 75: 1243-1252.

Miller, S.D., White, G.C., Sellers, R.A., Reynolds, H.V., Schoen, J.W., Titus, K., Barnes, V.G., Smith, R.B., Nelson, R.R., Ballard, W.B. and Schwartz, C.C. 1997. Brown and black bear density estimation in Alaska using radiotelemetry and replicated mark-resight techniques. Wildlife Monographs 133: 5-55.

Mowat, G. and Heard, D. C. 2006. Major components of grizzly bear diet across North America. Canadian Journal of Zoology 84: 473-489.

Mowat G, Heard, D.C., and Schwarz, C.J. 2013. Predicting grizzly bear density in western North America. PLoS ONE 8(12).

Mowat, G., Heard, D. C., Seip, D. R., Poole, K. G., Stenhouse, G. and Paetkau, D. W. 2005. Grizzly Ursus arctos and black bear U. americanus densities in the interior mountains of North America. Wildlife Biology 11: 31-48.

Nawaz, M. A. 2007. Status of the brown bear in Pakistan. Ursus 18: 89–100.

Nawaz. M.A., Swenson, J., and Zakaria, V. 2008. Pragmatic management increases a flagship species the Himalayan brown bears, in Pakistan’s Deosai National Park. Biological Conservation 141: 2230-2241.

Nielsen, S. E., Herrero, S., Boyce, M. S., Mace, R. D., Benn, B., Gibeau, M. L. and Jevons, S. 2004. Modelling the spatial distribution of human-caused grizzly bear mortalities in the Central Rockies ecosystem of Canada. Biological Conservation 120: 101-113.

Nielsen, S. E., Stenhouse, G. B. and Boyce, M. S. 2006. A habitat-based framework for grizzly bear conservation in Alberta. Biological Conservation 130: 217-229.

Proctor, M. F., Mclellan B. N., Strobeck, C. and Barclay, R. M. R. 2005. Genetic analysis reveals demographic fragmentation of grizzly bears yielding vulnerably small populations. Proceedings of the Royal Society of London B Biological Sciences 272: 2409-2416.

Proctor, M.F., Paetkau, D., McLellan, B.N., Stenhouse, G.B., Kendall, K.C., Mace, R.D., Kasworm, W.F., Servheen, C., Lausen, C.L., Gibeau, M.L., Wakkinen, W.L., Haroldson, M.A., Mowat, G., Apps, C.D., Ciarniello, L.M., Barclay, R.M.R., Boyce, M.S., Strobeck, C., and Schwartz, C.C. 2012. Population fragmentation and inter-ecosystem movements of grizzly bears in western Canada and the northern United States. Wildlife Monographs 180: 1-46.

Proctor, M. F., Servheen, C., Miller, S. D., Kasworm, W. F. and Wakkinen, W. L. 2004. A comparative analysis of management options for grizzly bear conservation in the US-Canada trans-border area. Ursus 15: 145-160.

Ridings, C. 2006. Green bear in the desert. International Bear News 15(2): 12-13.

Sacco, T. and Van Valkenburgh, B. 2004. Ecomorphological indicators of feeding behaviour in the bears (Carnivora: Ursidae). Journal of Zoology (London) 263: 41-54.

Sathyakumar, S. 2006. The status of brown bears in India. Understanding Asian bears to secure their future, pp. 7-11. Japan Bear Network, Ibaraki, Japan.

Sathyakumar S., Kaul, R., Ashraf, N.V., Mookerjee, A., and Menon, V. 2012. National bear conservation and welfare action plan. Ministry of Environment and Forests, Wildlife Institute of India and Wildlife Trust of India.

Schwartz, C.C.,Haroldson, M.A., G.C., Cherry, S., Keating, K.A., Moody, D., and Servheen, C. 2006. Temporal, spatial, and environmental influences on the demographics of grizzly bears in the greater Yellowstone Ecosystem. Wildlife Monographs 161(1).

Schwartz, C. C., Miller, S. D. and Haroldson, M. A. 2003. Grizzly bear. In: G. A. Feldhamer, B. C. Thompson, and J. A. Chapman (eds), Wild mammals of North America: biology, management, and conservation, pp. 556-586. Johns Hopkins University Press, Baltimore, Maryland, USA.

Servheen, C. 2013. Bear paw smuggling into China continues. International Bear News 22(3): 24.

Servheen, C., Herrero, S. and Peyton, B. 1999. Bears. Status Survey and Conservation Action Plan. IUCN/SSC Bear and Polar Bear Specialist Groups, Gland, Switzerland.

Seryodkin, I. 2006. The biology and conservation status of brown bears in the Russian Far East. Understanding Asian bears to secure their future, pp. 79-85. Japan Bear Network, Ibaraki, Japan.

Swenson, J.E., Sandergren, F., and Söderberg, A. 1998. Geographic expansion of an increasing brown bear population: evidence for presaturation dispersal. Journal of Animal Ecology 67: 819–82.

Swenson, J., Gerstl, N., Dahle, B. and Zedrosser, A. 2000. Action plan for the conservation of the brown bear in Europem (Ursus arctos). Council of Europe, Strasbourg, France.

Tumendemberel, O., Proctor, M., Reynolds, H., Boulanger, J., Luvsamjamba, A., Tserenbataa, T., Batmunkh, M., Craighead, D., Yanjin, N., Paetkau, D. 2015. Gobi bear abundance and inter-oases movement survey, Gobi Desert, Mongolia. Ursus 26: 129-142.

U.S. Fish and Wildlife Service. 2005. Endangered and threatened wildlife and plants; Designating the greater Yellowstone ecosystem population of grizzly bears as a distinct population segment; removing the Yellowstone distinct population segment of grizzly bears from the federal list of endangered and threatened wildlife. United States Federal Register. 70: 69854-69884.

Wakkinen, W. L. and Kasworm, W. F. 2004. Demographics and population trends of grizzly bears in the Cabinet-Yaak and Selkirk ecosystems of British Columbia, Idaho, Montana, and Washington. Ursus 15: 65-75.

Waller, J. S. and Servheen, C. 2005. Effects of transportation infrastructure on grizzly bears in northwestern Montana. Journal of Wildlife Management 69: 985-1000.

Wiegand, T., Stephan, T., Naves, J. and Fernandez, A. 1998. Assessing the risk of extinction for the brown bear (Ursus arctos) in the Cordillera Cantabrica, Spain. Ecological Monographs 68: 539-570.

Zedrosser, A., Dahle, B., Swenson, J. E. and Gerstl, N. 2001. Status and managment of the brown bear in Europe. Ursus 12: 9-20.

Citation: McLellan, B.N., Proctor, M.F., Huber, D. & Michel, S. 2017. Ursus arctos (amended version of 2017 assessment). The IUCN Red List of Threatened Species 2017: e.T41688A121229971. . Downloaded on 24 May 2018.
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