Pusa caspica


Bibliography [top]

Allchin, C., Barrett, T., Duck, C., Eybatov, T., Forsyth, M., Kennedy, S. and Wilson, S. 1997. Surveys of Caspian seals in the Apsheron Peninsula region and residue and pathology analyses of dead seal tissues. In: H. Dumont, S. Wilson and B. Wazniewicz (eds), Caspian Environment Programme, the First Bio-Network Workshop, November, 1997., pp. 101-118. Bordeaux, France.

Anan, Y., Kunito, T., Ikemoto, T., Kubota, R., Watanabe, I., Tanabe, S., Miyazaki, N. and Petrov, E. A. 2002. Elevated concentrations of trace elements in Caspian seals (Phoca caspica) found stranded during the mass mortality events in 2000. AArchives of Environmental Contamination and Toxicology 42: 354-362.

Arnason, A., Gullberg, A., Janke, A., Kullberg, M., Lehman, N., Petrov, E. A. and Väinölä, R. 2006. Pinniped phylogeny and a new hypothesis for their origin and dispersal. Molecular Phylogenetics Evolution 41: 345–354.

Badamshin, B. I. 1970. Resources of the Caspian seal and the means of its rational utilization. Translation by Naval Oceanographic Office Washington D.C. from: Badamshin, B.I. 1961. Zapay kaspiiskogo tyulenya i puti ikh ratsiona’lnogo ispol’zovaniya. Tr. Soveshch. Ikhitiol. Komis. AN SSSR.

Bininda-Emonds, O. R. P., Cardillo, M., Jones, K. E., MacPhee, R. D. E., Beck, R. M. D., Grenyer, R., Price, S. A., Vos, R. A., Gittleman, J. L. and Purvis, A. 2007. The delayed rise of present day mammals. Nature 446: 507–512.

Caspian Environment Programme. 2007. Caspian Seal conservation Action Plan. Available at:

Dumont, H. J. 1998. The Caspian Lake: history, biota, structure, and function. Limnology Oceanogaphy 43: 44-52.

Eybatov, T., Asadi, H., Erokhin, P., Kuiken, T., Jepson, P., Deaville, R. and Wilson, S. 2002. Caspian seal mortality. Ecotox Final Report, Appendix A2. World Bank.

Frost, K. J. and Lowry, L. F. 1981. Ringed, Baikal and Caspian seals Phoca hispida Schreber, 1775; Phoca sibirica Gmelin, 1788; and Phoca caspica Gmelin, 1788. In: S. H. Ridgway and R. Harrison (eds), Handbook of marine mammals, Vol. 2: Seals, pp. 29-53. Academic Press.

Härkönen, T., Jüssi, M., Baimukanov, M., Bignert, A., Dmitrieva, L., Kasimbekov, Y., Verevkin, M., Wilson, S. and Goodman, S.J. 2008. Pup production and breeding distribution of the Caspian Seal (Phoca caspica) in relation to human impacts. Ambio 37(5): 356-361.

Härkönen, T., Jüssi, M., Baimukanov, M., Dmitrieva, L., Kasimbekov, Y., Verevkin, M., Wilson, S. and Goodman, S. 2005. Population size and density distribution of the Caspian seal (Phoca caspica) on the winter ice field in Kazakh waters 2005. Available at:

Heptner, V. G., Chapskii, K. K., Arsen’ev, V. A. and Sokolov, V. E. 1996. Mammals of the Soviet Union. Smithsonian Institution Libraries and National Science Foundation.

IUCN. 2008. 2008 IUCN Red List of Threatened Species. Available at: (Accessed: 5 October 2008).

Ivanov, V. P., Kamakin, A. M., Ushivitzev, V. B., Shiganova, T., Zhukova, O., Aladin, N., Wilson, S. C., Harbison, G. R. and Dumont, H. 2000. Invasion of the Caspian Sea by the comb jellyfish Mnemiopsis leidyi (Ctenophora). Biological Invasions 2: 255-258.

Kajiwara, N., Niimi, S., Watanabe, M., Ito, Y., Takahashi, S., Tanabe, S., Khuraskin, L. S. and Miyazaki, N. 2002. Organochlorine and organotin compounds in Caspian seals (Phoca caspica) collected during the unusual mortality event in the Caspian Sea in 2000. Environmental Pollution 117: 391-402.

Kennedy, S., Kuiken, T., Jepson, P. D., Deaville, R., Forsythe, M., Barrett, T., van de Bildt, M. W. G., Osterhaus, A. D. M. E., Eybatov, E., Duck, C., Kydyrmanov, A., Mitrofanov, I. and Wilson, S. 2000. Mass die-off of Caspian seals caused by canine distemper virus. Emerging Infectious Diseases 6: 637-639.

Khuraskin, L. S. and Zakharova, N. A. 2002. Phoca (Pusa) caspica Gmelin, 1788. Available at:

Kideys, A. E., Roohi, A., Bagheri, S., Finenko, G. and Kamburska, L. 2005. Impacts of invasive ctenophores on the fisheries of the Black Sea and Caspian Sea. Oceanography 18: 76-85.

Kosarev, A. N. and Yablonskaya, E. A. 1994. The Caspian Sea. SPB Academic Publishing, The Hague, The Netherlands.

Krylov, V. I. 1990. Ecology of the Caspian seal. Finnish Game Reserve 47: 32-36.

Kuiken, T., Kennedy, S., Barrett, T., Borgsteede, F. H., Van de Bildt, M. W. G., Brew, S. D., Codd, G. A., Duck, C., Deaville , R., Eybatov, T., Forsyth, M., Foster, G., Jepson, P., Kydyrmanov, A., Mitrofanov, I., Ward, C. J., Wilson, S. and Osterhaus, A. D. M. E. 2006. The 2000 canine distemper epidemic in Caspian seals (Phoca caspica): pathology and analysis of contributory factors. Veterinary Pathology 43: 321-338.

Miyazaki, N. 2002. Ringed, Caspian, and Baikal seals Pusa hispida, P. caspica, and P. sibirica. In: W. F. Perrin, B. Wursig and J. G. M. Thewissen (eds), Encyclopedia of Marine Mammals, pp. 1033-1037. Academic Press.

Palo, J. U. and Väinöla, R. 2006. The enigma of the land-locked Baikal and Caspian seals addressed through phylogeny of phocine mitochondrial sequences. Biological Journal of the Linnean Society 88: 61-72.

Piletskii, A. M. and Krylov, V. I. 1990. Caspian seals of the Volga Delta. In: V. I. Krylov (ed.), Some remarks to biology and ecology of the Caspian seal, pp. 58-78. VNIRO, Moscow, Russia.

Popov, L. 1979. Caspian Seal. Mammals in the Seas, Vol. II: Pinniped species summaries and report on sirenians, pp. 74-75.

Popov, L. 1982. Status of the main ice-living seals inhabiting inland waters and coastal marine areas of the USSR. Mammals in the Seas, Vol. IV: Small cetaceans, sirenians, seals and otters., pp. 361-381.

Reijnders, P., Brasseur, S., van der Toorn, J., van der Wolf, P., Boyd, I., Harwood, J., Lavigne, D. and Lowry, L. 1993. Seals, fur seals, sea lions, and walrus. Status survey and conservation action plan. IUCN Seal Specialist Group.

Rice, D.W. 1998. Marine Mammals of the World. Systematics and Distribution. Society for Marine Mammalogy, Lawrence, Kansas.

St. Aubin, D. J. 1990. Physiologic and toxic effects on pinnipeds. In: J. R. Geraci and D. J. St. Aubin (eds), Sea mammals and oil: confronting the risks, pp. 103-127. Academic Press, New York, USA.

Watanabe, M., Tanabe, S., Tatsukawa, R., Amano, M., Miyazaki, N., Petrov, E. A. and Khuraskin, S. L. 1999. Contamination levels and specific accumulation of persistent organochlorines in Caspian seal (Phoca caspica) from the Caspain Sea, Russia. Archives of Environmental Contamination and Toxicology 37: 396-407.

Citation: Härkönen, T. (IUCN SSC Pinniped Specialist Group) 2008. Pusa caspica. The IUCN Red List of Threatened Species. Version 2014.3. <>. Downloaded on 20 December 2014.
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