Spheniscus humboldti

Ayala, L.; Sánchez, R.; Kelez, S.; and Vásquez, F. 2007. Estudio poblacional del Pingüino de Humboldt Spheniscus humboldti en la costa centro y sur del Perú en el invierno del 2004.

Boersma, P. D.; Redstock, G. A.; Stokes, D. L.; Majluf P. 2006. Oceans apart: conservation models for two temperate penguin species shaped by the marine environment. Marine Ecology Progress Series 335: 217-225.

Cheney, C. 1998. The current situation of the Humbolt Penguin in Chile and Peru: a report from the population and habitat viability analysis meeting, part 1. Penguin Conservation 11: 4-9.

Culik, B. M.; Luna-Jorquera, G. 1997. Satellite tracking of Humboldt penguins (Spheniscus humboldti) in northern Chile. Marine Biology 128: 547-556.

Ellis, S.; Croxall, J. P.; Cooper, J. 1998. Penguin conservation assessment and management plan: report from the workshop held 8-9 September 1996, Cape Town, South Africa. IUCN/SSC, Apple Valley, USA.

Hays, C. 1984. The Humboldt Penguin in Peru. Oryx 18: 92-95.

Hays, C. 1986. Effects of the 1982-83 El Niño on Humboldt Penguin colonies in Peru. Biological Conservation 36: 169-180.

IUCN. 2012. IUCN Red List of Threatened Species (ver. 2012.1). Available at: http://www.iucnredlist.org. (Accessed: 19 June 2012).

Luna-Jorquera, G. and Culik, B.M. 1999. Diving behaviour of Humboldt penguins Spheniscus humboldti in northern Chile. Marine Ornithology 27: 67-76.

Morales Sanchez, J. E. 1988. Confirmación de la presencia de Spheniscus humboldti Meyen (Aves: Spheniscidae) Para Colombia. Trianea (Acta Cientifica y Tecnologia INDERENA): 141-143.

Paredes, R.; Zavalaga, C. B. 2001. Nesting sites and nest types as important factors for the conservation of Humboldt Penguins (Sphensicus humboldti). Biological Conservation 100: 199-205.

Paredes, R.; Zavalaga, C. B.; Battistini, G.; Majluf, P.; McGill, P. 2003. Status of the Humboldt Penguin in Peru, 1999-2000. Waterbirds 26: 129-138.

Taylor S.S; Leonard M.L; Boness D.J. 2001. Foraging trip duration increases for Humboldt Penguins tagged with recording devices. Journal of Avian Biology 32(4): 369-372.

Taylor, SS ; Leonard, ML ; Boness, DJ ; Majluf, P. 2002. Foraging by Humboldt penguins (Spheniscus humboldti) during the chick-rearing period: general patterns, sex differences, and recommendations to reduce incidental catches in fishing nets . Canadian Journal Of Zoology-Revue Canadienne De Zoologie 80(4): 700-707.

Taylor, S.S.; Leonard, M. L.; Boness, D.J.; Majluf, P. 2004. Humboldt Penguins Spheniscus humboldti change their foraging behaviour following breeding failure. Marine Ornithology 32: 63-67.

Vilina, Y. A.; Capella, J. J.; González, J.; Gibbons, J. E. 1995. Apuntes para la conservación de las aves de la reserva nacional Pingüino de Humboldt. Boletín Chileno de Ornitología: 2-6.

Wallace, R. S.; Grzybowski, K.; Diebold, E.; Michaels, M. G.; Tear, J. A.; Willis, M. J. 1999. Movements of Humboldt Penguins from a breeding colony in Chile. Waterbirds 22: 441-444.

Williams, T. D. 1995. The penguins Spheniscidae. Oxford University Press, Oxford.